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The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and Spirula. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name.
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
Actinoceras is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician. It is an extinct genus of nautiloid cephalopod that thrived in the warm waters of the United States during the Paleozoic era.
Ormoceras is an actinocerid nautiloid genus and type for the family Ormoceratidae, found in North America from the late Chazyan through the early Cincinnatian of the Middle and Upper Ordovician, but which continued through the Devonian worldwide.
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian, in which the connecting rings are thin and siphuncle segments are variably expanded. At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three.
The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.
Winnipegoceras is an extinct nautiloid genus from the Ordovician belonging to the Order Discosorida.
Dawsonoceratidae is an extinct family of orthoconic nautiloid cephalopods that lived in what would be North America and Europe from the Late Ordovician through the Middle Devonian from about 480–390 mya, existing for approximately 90 million years.
Sactoceras is an extinct nautiloid cephalopod that lived during the Ordovician and Silurian in what would become North America, Europe, and Asia.
Carbactinoceratidae is a family of extinct cephalopods with external shells that lived around 325 million years ago, during the Carboniferous period. They were the last group of orthocones to attain sizes exceeding one meter in length. One specimen of the carbactinoceratid species Rayonnoceras solidiforme, recently found in Arkansas, was measured at 2.5 metres (8.2 ft) long.
Oncoceratidae is a family of nauatiloid cephalopods in the order Oncocerida established by Hyatt, 1884, that range from the Middle Ordovician to the Upper Silurian.
The Armenoceratidae are a family of early Paleozoic nautiloid cephalopods belonging to the order Actinocerida.
Wutinoceras is a genus of now extinct nautiloid cephalopods of the Wutinoceratidae family. It exhibits orthoconic actinocerids with ventral siphuncles composed of broadly expanded segments.
Macroloxoceras is a large pseuorthocerid from the upper Devonian of Central Colorado and Southern New Mexico with features resembling those found in actinocerids. Pseudorthocerids and actinocerids are extinct nautiloid cephalopods, generally with long straight shells and expanded siphuncle segments filled with organic deposits.
Landeroceras is a genus of straight shelled cyrtogomphoceratid from the Middle Ordovician Big Horn dolomite of Wyoming.
Poterioceratidae is a family of nautiloid cephalopods included in the Oncocerida that lived during the period from the Early Devonian to the Early Carboniferous (Mississippian). Members of the Poterioceratidae are distinguished by a subcircular to compressed exogastric shell that has no hyponomic sinus and a central to subcentral siphuncle composed of subquadrate to nummuloidal segments in which the septal necks are more strongly curved on the upper, or dorsal side. This is opposite from the Karoceratidae in which siphuncle segments are inflated ventrally but straight dorsally. Some poterioceratid genera have actinosiphonate structures or annular deposits within the siphuncle. In others it is empty.
The Sactorthoceratidae comprise Orthocerataceaen genera with a subcentral suborthochoanitic siphuncle composed of slightly expanded segments and free of organic deposits. The camerae (chambers) of the phragmocone likewise have organic deposits that are typically retarded or sparse.
The Barrandeocerina comprise a suborder of Early Paleozoic nautiloid cephalopods, primitively coiled but later forms may be cyroconic, gyroconic, torticonic, and even breviconic, all having empty siphuncles with thin connecting rings. The Barrandeocerina were originally defined as a separate order by Rousseau Flower, but since then have been united within the Tarphycerida as a suborder. Derivation is from the Tarphyceratidae.
Discosoridae comprise a family of endogastric discosorids,, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.
Karoceratidae is a family of nautiloids within the order Oncocerida, characterized by straight or curved, laterally narrow, shells and slender, ventral siphuncles that are empty except in Karoceras. Siphuncle segments are inflated ventrally but straight dorsally. Septal necks are cyrtochoanitic, outwardly curved; connecting rings are thin.
References
- ↑ Walter C. Sweet (1964). "Nautiloidea- Orthocerida". In Raymond C. Moore (ed.). Treatise on Invertebrate Paleontology, Part K. Geological Society of America and University of Kansas Press. p. K246.
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