Pseudorthocerida

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Pseudorthocerida
Temporal range: Silurian–Triassic [1]
Reticycloceras sp Carboniferous Bear Gulch Montana USA Mo1832b.jpg
Reticycloceras , a Carboniferous pseudorthocerid
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Orthoceratoidea
Order: Pseudorthocerida
Barskov 1963, from Flower & Caster, 1935

Pseudorthocerida is an order of generally straight longiconic orthoceratoids with a subcentral to marginal cyrtochoanitic siphuncle composed of variably expanded segments which may contain internal deposits that may develop into a continuous parietal lining. (Sweet 1964 [2] ). Cameral deposits are common and concentrated ventrally. Apices typically have a slight to moderate exogastric curvature

Contents

The Pseudorthocerida are included in a broad in-group of generally orthoconic cephalopods known as the Orthoceratoidea (Kroger 2008) along with the Ascocerida, Dissidocerida, Lituitida, and Orthocerida.

The Pseudorthocerida were among the last living orthoconic nautiloids. One family, the Trematoceratidae, survived into the Triassic Period.

Taxonomy

Current understanding

By current understanding the Pseudorthocerida contains the following families.

Pseudorthoceratidae
Cayutoceratidae
Pseudactinoceratidae
Spyroceratidae
Carbactinoceratidae
Trematoceratidae [1]

The Pseudorthoceratidae through Spyroceratidae are presented as subfamilies in Sweet (1964) and are included in the Pseudorthoceratidae sensu Sweet (1964). The Carbactinoceratidae are removed from the Actinocerida in Kroger and Mapes (2007).

Earlier perspectives

Pseudorthocerids were previously known as the Pseudorthocerataceae, a superfamily within the Orthocerida (Sweet 1964) which included the Silurian and later Pseudorthoceratidae and Middle Ordovician to Middle Silurian Proteoceratidae. Both have siphuncles that are partly cyrochoanitic with expanded segments and internal deposits but differ in the manner in which they develop.

Flower (1976) distinguished Middle Ordovician pseudorthocerids from Silurian and later pseudorthocerids on the basis of how the siphuncle changes during the life of the animal. The earlier Proteoceratidae begin cyrtochoanitic with expanded segments and end up orthochoanitic with subcylindrical segments. As juveniles they are "pseudorthoceratoid", as mature individuals they are "orthoceratoid". In contrast the later Pseudorthoceratidae, begin orthochoanitic with subcylindrical segments and ontogenetically become cyrtochoanitic with expanded segments. As such they matured into "pseudorthoceratoids" from "orthoceratoid" juveniles. From this Flower (1976) concluded that the Ordovician and Silurian pseudorthocerids (sensu Sweet 1964) were derived from different orthocerids (Orthocerida) and were therefore polyphyletic. As a result, true pseudorthocerids were limited to the Pseudorthoceratidae while the earlier Ordovician look-a-likes were retained in the Orthocerida (Michelinocerida Flower)

Related Research Articles

Nautiloid Extant subclass of cephalopods

Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day.

Orthoceratidae Extinct family of molluscs

Orthoceratidae is an extinct family of actively mobile carnivorous cephalopods, subclass Nautiloidea, that lived in what would be North America, Europe, Asia, Africa, and Australia from the Ordovician through Triassic from 490—203.7 mya, existing for approximately 286.4 million years.

Orthocerida Extinct order of molluscs

Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician possibly to the Late Triassic. A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous. They were most common however from the Ordovician to the Devonian.

Bactritida Fossil order of cephalopods

The Bactritida are a small order of more or less straight-shelled (orthoconic) cephalopods that first appeared during the Emsian stage of the Devonian period with questionable origins in Pragian stage before 409 million years ago, and persisted until Carnian pluvial event in the upper middle Carnian stage of the Triassic period. They are considered ancestors of the ammonoids, as well as of the coleoids.

Ascocerida Extinct order of molluscs

The Ascocerida are comparatively small, bizarre Orthoceratoids known only from Ordovician and Silurian sediments in Europe and North America, uniquely characterized by a deciduous conch consisting of a longiconic juvenile portion and an inflated breviconic adult portion that separate sometime in maturity.

Oncocerida Extinct order of nautiloids

The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian, in which the connecting rings are thin and siphuncle segments are variably expanded. At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three.

The Actinocerida are an order of generally straight, medium to large cephalopods that lived during the early and middle Paleozoic, distinguished by a siphuncle composed of expanded segments that extend into the adjacent chambers, in which deposits formed within contain a system of radial canals and a narrow space along the inner side of the connecting ring known as a paraspatium. Septal necks are generally short and cyrtochoanitic, some being recumbent, some hook shaped. Most grew to lengths of about 60 to 90 cm but some, like the Huroniidae of the Silurian grew significantly larger.

Pseudactinoceras is an extinct nautiloid cephalopod included in the order Pseudorthocerida and the namesake of the family Pseudactinoceratidae.

<i>Kionoceras</i> Extinct genus of nautiloids

Kionoceras is an extinct nautiloid cephalopod genus included in the orthocerid family Kionoceratidae with scattered worldwide distribution from the Middle Ordovician to the Lower Permian. Kionoceratids are orthocerids with prominent longitudinal ornamentation on their shells, sometimes augmented by secondary transverse ornamentation. Orthocerids are, of course, prehistoric nautiloides with generally straight and elongate shells, mostly with central or subcentral siphuncles.

Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.

Geisonoceratidae Extinct family of molluscs

Geisonoceratidae is an extinct family of orthoceroid cephalopods endemic to what would be Asia, Europe, and North America from the Middle Ordovician to the Middle Devonian living from about 470—380 mya, existing for approximately 90 million years. With the possible addition of an Early Cretaceous orthocerid from the western Caucasus the range of this group increases dramatically to some 350 million years, thus making it one of the longest lived families of the Nautiloidea.

Rioceras is an extinct genus of orthocerid cephalopods of the family Baltoceratidae that lived in what would be North America during the Early Ordovician from 480—472 mya, existing for approximately 8 million years.

Sactoceras is an extinct nautiloid cephalopod that lived during the Ordovician and Silurian in what would become North America, Europe, and Asia.

Orthoceratoidea Extinct subclass of cephalopods

Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars, extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer.

Phragmoceratidae Extinct family of molluscs

The Phragmoceratidae is a family of extinct nautiloid cephalopods from the Order Discosorida that lived during the latter part of the Silurian.

The Orthocerataceae is a superfamily of orthocerid cephalopods that lived from the late Early Ordovician to the Early Cretaceous, but is no longer in general use.

Kionoceratidae Family of nautiloids

Kionoceratidae is a family in the Orthocerida, proposed by Hyatt in 1900 for genera characterized by prominent ornamentation in the form of longitudinal ribs, ridges, or lirae, or combinations thereof, sometimes with similar transverse ornament or faint transverse annulations.

Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.

Murrayoceras is a nautilid cephalopod included in the orthocerid family Baltoceratidae, widespread in the Middle Ordovician of North America, characterized by a depressed orthoconic shell with a subtriangular cross section and flattened venter and a proportionally large ventral siphuncle, 0.15 to 0.3 the dorso-ventral shell diameter. Septa are close spaced with sutures forming broad lobes on the upper flanks and ventral surface.

Ascoceratidae is a family of Ascocerida, bizarre orthoceratoid cephalopods, with longiconic, deciduous early growth stages which undergo period truncation and an inflated breviconic mature ascoceroid stage. The Ascoceratidae are divided into two, unequal subfamilies.

References

  1. 1 2 Zakharov, Yuri D. (1996). "Orthocerid & ammonoid shell structure: its bearing on cephalopod classification". Bulletin of the National Science Museum. Series C, Geology & Paleontology. 22 (1, 2): 11–35.
  2. Sweet, Walter C. (1964). Nautiloidea -- Orthocerida, in Treatise on Invertebrate Paleontology. Part K. Mollusca 3. Geological Society of America, and University of Kansas Press, New York, New York and Lawrence, Kansas.