Carbactinoceratidae Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Order: | † Pseudorthocerida |
Family: | † Carbactinoceratidae Schindewolf, 1943 |
Genera [2] | |
Carbactinoceratidae is a family of extinct cephalopods with external shells that lived around 325 million years ago, during the Carboniferous period. They were the last group of orthocones to attain sizes exceeding one meter (three feet) in length. One specimen of the carbactinoceratid species Rayonnoceras solidiforme, recently found in Arkansas, was measured at 2.5 metres (8.2 ft) long.
Shells of carbactinoceratids are usually straight or slightly curved, with a large central or subcentral siphuncle. The first chamber at the apex is long and conical, the first septum slightly convex with the a septal perforation about one third of the shell diameter and a cyrtochoanitic septal neck (tubular or funnel-like projection from the back of the septum. The first siphuncle segment is elongated, subsequent ones formed during growth of the animal are subglobular (rounded but not spherical). Organic deposits are formed within the siphuncle through which run central, and radial canals along with a seam along the connecting rings known as a perispatium. [3]
Schindewolf (1941) and Teichert (1964) classified the Carbactinoceratidae within the actinocerids based on siphuncle similarity. [4] In both groups the siphuncle contains collar-like deposits and radial canals. [3]
However, there are some problems with the actinocerid hypothesis. Carbactinocerids such as Carbactinoceras appear much later than the last definite actinocerids, leaving a large stratigraphic gap between the two. [5] Furthermore, carbactinoceratids share key characters from the early development of pseudorthocerids missing from actinocerids. Among these is a bullet-shaped apex that is not constricted, and a first segment of the siphuncle that is relatively narrow. [3] Moreover, pseudorthocerids abounded in the Carboniferous period and so eliminate the stratigraphic gap.
Rayonnoceras is a genus of extinct cephalopods that lived around 325 million years ago during the Carboniferous. Although they resemble earlier actinocerids they are now thought to belong to the Pseudorthocerida
The Bactritida are a small order of more or less straight-shelled (orthoconic) cephalopods that first appeared during the Emsian stage of the Devonian period with questionable origins in the Pragian stage before 409 million years ago, and persisted until the Carnian pluvial event in the upper middle Carnian stage of the Triassic period. They are considered ancestors of the ammonoids, as well as of the coleoids.
Huronia is an actinocerid genus included in the Huroniidae along with Discoactinoceras and Huroniella,(Teichert 1964). Huronia is characterized by long siphuncle segments with the free part of the connecting rings only slightly inflated and by a narrow central canal and strongly curved radial canals located in the anterior part of each siphuncle segment
Actinoceras is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician. It is an extinct genus of nautiloid cephalopod that thrived in the warm waters of the United States and England during the Paleozoic era.
Ormoceras is an actinocerid nautiloid genus and type for the family Ormoceratidae, found in North America from the late Chazyan through the early Cincinnatian of the Middle and Upper Ordovician, but which continued through the Devonian worldwide.
Lambeoceras is a genus of large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America and the sole representative of the family Lambeoceratidae.
Armenoceras is a genus of actinocerid nautiloid cephalopods whose fossils ranged from the late Whiterockian Stage in the early Middle Ordovician, through the remainder of the period and on into the Upper Silurian. It is the type genus of the family Armenoceratidae.
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian, in which the connecting rings are thin and siphuncle segments are variably expanded. At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three.
Cyrtonybyoceras is a genus of slightly exogastric members of the Wutinoceratidae, a family of actinocerids and probably derived from an earlier Wutinoceras. The shell of Cyrtonybyoceras is curved slightly upwardly and is slightly compressed. Sutures slope toward the aperture, from the dorsum to the venter. The siphuncle is ventral but not marginal, in general form like that of Nybyoceras. Upper and lower septal necks are recumbent or narrowly free. The canal system is reticular, characteristic of the Wutinoceratidae.
The Actinocerida are an order of generally straight, medium to large cephalopods that lived during the early and middle Paleozoic, distinguished by a siphuncle composed of expanded segments that extend into the adjacent chambers, in which deposits formed within contain a system of radial canals and a narrow space along the inner side of the connecting ring known as a paraspatium. Septal necks are generally short and cyrtochoanitic, some being recumbent, some hook shaped. Most grew to lengths of about 60 to 90 cm but some, like the Huroniidae of the Silurian grew significantly larger.
The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.
Paraloxoceras is a genus of straight shelled, orthoconic nautiloid cephalopods, now extinct, that lived during the Early Carboniferous. Fossils have been found in Europe and central Asia; the type, P. konincki, named by Flower, came from Belgium.
Winnipegoceras is an extinct nautiloid genus from the Ordovician belonging to the Order Discosorida.
The Actinoceriatidae are a family of actinocerids named by Saemann in 1853 for those that grew to have large shells with blunt apices and large siphuncles with widely expanded segments and a generally arcuate endosiphucular canal system. Their range is from the upper Middle Ordovician to the Lower Silurian. Actinocerids are generally straight-shelled nautiloid cephalopods with a siphuncle composed of expanded segments, typically with thin connecting rings, in which the internal deposits are penetrated by a system of canals
The Armenoceratidae are a family of early Paleozoic nautiloid cephalopods belonging to the order Actinocerida.
Nybyoceras is an actinocerid genus assigned to the Armenoceratidae and similar to Armenoceras except for having a siphuncle close to the ventral side of the shell.
Macroloxoceras is a large pseuorthocerid from the upper Devonian of Central Colorado and Southern New Mexico with features resembling those found in actinocerids. Pseudorthocerids and actinocerids are extinct nautiloid cephalopods, generally with long straight shells and expanded siphuncle segments filled with organic deposits.
Kobyashiceras is an extinct genus in the cephalopod order Actinocerida, from Lower Devonian marine sediments in Japan. The type and sole included species is Kobayashiceras gifuense. The generic name honors the late Dr. Teiichi Kobayashi who contributed greatly to the study of Paleozoic cephalopods. The specific name is derived from Gifu, the name of the prefecture in which the type locality is found.
Poterioceratidae is a family of nautiloid cephalopods included in the Oncocerida that lived during the period from the Early Devonian to the Early Carboniferous (Mississippian). Members of the Poterioceratidae are distinguished by a subcircular to compressed exogastric shell that has no hyponomic sinus and a central to subcentral siphuncle composed of subquadrate to nummuloidal segments in which the septal necks are more strongly curved on the upper, or dorsal side. This is opposite from the Karoceratidae in which siphuncle segments are inflated ventrally but straight dorsally. Some poterioceratid genera have actinosiphonate structures or annular deposits within the siphuncle. In others it is empty.
Carbactinoceras is a genus of Early Carboniferous (Visean) actincoceroids first found in Europe (Germany) related to Rayonnoceras but probably smaller.