| Keuppia Temporal range: Late Cenomanian | |
|---|---|
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| K. levante | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Mollusca |
| Class: | Cephalopoda |
| Order: | Octopoda |
| Family: | † Palaeoctopodidae |
| Genus: | † Keuppia Fuchs, Bracchi & Weis, 2009 |
| Species | |
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Keuppia is an extinct genus of octopus.
It consists of two species, Keuppia hyperbolaris and Keuppia levante, both of which lived approximately 95 million years ago. [1] Both species were found in fossilized form, which is very uncommon for extinct octopuses, as the soft tissue of dead octopuses almost always disintegrates before it has a chance to fossilize. [2] These fossils, along with those of the genus Styletoctopus , were found from the Cretaceous-age Hâqel and Hjoula localities in Lebanon. The presence of a gladius vestige in this genus shows a transition from squid to octopus in which the inner shell has divided in two in early forms to eventually be reduced to lateralized stylets, as can be seen in Styletoctopus.
Ammonoids are extinct spiral shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living coleoids than they are to shelled nautiloids. The earliest ammonoids appeared during the Devonian, with the last species vanishing during or soon after the Cretaceous–Paleogene extinction event. They are often called ammonites, which is most frequently used for members of the order Ammonitida, the only living group of ammonoids from the Jurassic up until their extinction.
Belemnoids are an extinct group of marine cephalopod, very similar in many ways to the modern squid. Like them, the belemnoids possessed an ink sac, but, unlike the squid, they possessed ten arms of roughly equal length, and no tentacles. The name "belemnoid" comes from the Greek word βbelemnon meaning "a dart or arrow" and the Greek word είδος, eidos meaning "form".
Cirrina or Cirrata is a suborder and one of the two main divisions of octopuses. Cirrate octopuses have a small, internal shell and two fins on their head, while their sister suborder Incirrina has neither. The fins of cirrate octopods are associated with a unique cartilage-like shell in a shell sac. In cross-section, the fins have distinct proximal and distal regions, both of which are covered by a thin surface sheath of muscle.
Protosphyraena is a fossil genus of swordfish-like marine fish, that thrived worldwide during the Upper Cretaceous Period (Coniacian-Maastrichtian). Though fossil remains of this taxon have been found in both Europe and Asia, it is perhaps best known from the Smoky Hill Member of the Niobrara Chalk Formation of Kansas. Protosphyraena was a large fish, averaging 2–3 metres in length. Protosphyraena shared the Cretaceous oceans with aquatic reptiles, such as mosasaurs and plesiosaurs, as well as with many other species of extinct predatory fish. The name Protosphyraena is a combination of the Greek word protos ("early") plus Sphyraena, the genus name for barracuda, as paleontologists initially mistook Protosphyraena for an ancestral barracuda. Recent research shows that the genus Protosphyraena is not at all related to the true swordfish-family Xiphiidae, but belongs to the extinct family Pachycormidae.
Palaeoctopus is an extinct genus of octopuses that lived during the Late Cretaceous. It contains one valid species, P. newboldi, which has been found in Lebanon.
Eupodophis is an extinct genus of snake from the Late Cretaceous period. It has two small hind legs and is considered a transitional form between Cretaceous lizards and limbless snakes. The feature, described as vestigial, was most likely useless to Eupodophis. The type species Eupodophis descouensi was named in 2000 and resides now in the paleontology section of the Mim Museum in Beirut, Lebanon. The specific name is dedicated to the French naturalist Didier Descouens.
Cearachelys is an extinct genus of pleurodiran turtle which existed some 110 million years ago. The genus is monotypic, with only type species Cearachelys placidoi known.
Paleontology or palaeontology is the study of prehistoric life forms on Earth through the examination of plant and animal fossils. This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2009.
Santanichthys diasii is a species of extinct fish that existed around 115 million years ago during the Albian age. S. diasii is regarded as the basal-most characiform, and is the earliest known member of Otophysi. It appears as a small fish, similar in appearance to a modern-day herring little more than 30 millimeters in length. Its most striking characteristic is the presence of a Weberian apparatus, which makes it the most primitive known member of the order Characiformes, the order in which modern-day tetras are classified. Santanichthys has been unearthed from numerous locations throughout Brazil, in rocks dating to the Cretaceous Period. Its presence in these strata is seen as an indicator for the biogeography and evolution of its order.
Coccodus is an extinct genus of marine pycnodontid fish that lived during the Late Cretaceous. The various species had a pair of massive, curved spines emanating from the lower sides of the head, and one curved spine on the top of its head. Unlike most pycnodontids, Coccodus species had a comparatively long body, giving the living animals a superficial resemblance to a scaly chimaera.
Styletoctopus is an extinct genus of octopus. The genus consists of the single species Styletoctopus annae, which lived approximately 95 million years ago during the late Cenomanian,. It was first discovered in 2009 by a team led by Dirk Fuchs of Freie University in the Hâqel and Hjoula localities in Lebanon. Very few octopus species appear in the fossil record, as octopuses consist of soft tissue that usually decomposes before it has time to fossilize.
The paleontological sites of Lebanon contain deposits of well preserved fossils and include some species found nowhere else. Notable among these is the Lebanese lagerstätten of the Late Cretaceous age, which contain a well-preserved variety of different fossils. Some fossils date back to the Jurassic period, and younger fossils of mammals from a different site belong to the Miocene through the Pleistocene.
Susisuchus is an extinct genus of neosuchian mesoeucrocodylian crocodyliform from the Early Cretaceous of Brazil. Fossils have been found from the Nova Olinda Member of the Aptian-age Crato Formation in the Araripe and Lima Campos Basins of northeastern Brazil. Named in 2003, Susisuchus is the sole member of the family Susisuchidae, and is closely related to the clade Eusuchia, which includes living crocodilians. The type species is S. anatoceps, known from a single partial articulated skeleton that preserves some soft tissue. A second species, S. jaguaribensis, was named in 2009 from fragmentary remains.
Belemnitida is an extinct order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous. Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most to the tip: the tongue-shaped pro-ostracum, the conical phragmocone, and the pointy guard. The calcitic guard is the most common belemnite remain. Belemnites, in life, are thought to have had 10 hooked arms and a pair of fins on the guard. The chitinous hooks were usually no bigger than 5 mm (0.20 in), though a belemnite could have had between 100 and 800 hooks in total, using them to stab and hold onto prey.
Lebanoraphidia is an extinct genus of snakefly in the family Mesoraphidiidae. The genus is solely known from Cretaceous, Upper Neocomian, fossil amber found in Lebanon. Currently the genus is composed of a single species Lebanoraphidia nana.
Pappotherium is an extinct genus of mammals from the Albian of Texas, US, known from a fossilized maxilla fragment bearing two tribosphenic molars, discovered within the Glen Rose Formation near Decatur, Wise County, Texas.
Mimodactylus is a genus of istiodactyliform pterosaur that lived in what is now Lebanon during the Late Cretaceous, 95 million years ago. The only known specimen was discovered in a limestone quarry near the town of Hjoula, belonging to the Sannine Formation. The owner of the quarry allowed the specimen to be prepared and scientifically described by an international team of researchers, and when it was eventually sold, the buyer donated it to the MIM Museum in Beirut. In 2019, the researchers named the new genus and species Mimodactylus libanensis; the generic name refers to the MIM Museum, combined with the Greek word daktylos for "digit", and the specific name refers to Lebanon. The well-preserved holotype specimen is the first complete pterosaur from the Afro-Arabian continent, and the third pterosaur fossil known from Lebanon.
Erquelinnesia is an extinct turtle genus from the Cretaceous period, which has been found in fossils. Paleontologist Louis Dollo first described this genus late 19th century from fossilized samples found in the layers of Upper Paleocene / Lower Eocene in Hainaut, Belgium. Erquelinnesia lived between the Upper Cretaceous and Lower Paleocene. Louis Dollo named his discovery as Pachyrhynchus gosseleti in 1886. However, in the following year, the nomenclature was changed by Dollo to Erquelinnesia gosseleti to resolve a conflict with another already existing generic name. Moreover, an additional species was attributed to the genus Erquelinnesia. The North American species called Erquelinnesia molaria that was known only from a fossilized mandible bone has subsequently re-categorized to the genus Euclastes.
Carolozittelia is an extinct genus of mammals, belonging to the order Pyrotheria. It contains the single species Carolozittelia tapiroides which lived during the Early Eocene. Its fossilized remains were found in South America.
Pseudastacus is an extinct genus of decapod crustaceans that lived during the Jurassic period in Europe, and possibly the Cretaceous period in Lebanon. Many species have been assigned to it, though the placement of some species remains uncertain and others have been reassigned to different genera. Fossils attributable to this genus were first described by Georg zu Münster in 1839 under the name Bolina pustulosa, but the generic name was changed in 1861 after Albert Oppel noted that it was preoccupied. The genus has been placed into different families by numerous authors, historically being assigned to Nephropidae or Protastacidae. Currently, it is believed to be a member of Stenochiridae.
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