Phragmoteuthida | |
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Comparation of hard parts of Phragmoteuthida, Belemnitida and Diplobelida | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Superorder: | † Belemnoidea |
Order: | † Phragmoteuthida Jeletzky in Sweet, 1964 |
Families | |
See text. |
Phragmoteuthida is an order of extinct coleoid cephalopods [1] characterized by a fan-like teuthoid pro-ostracum attached to a belemnoid-like phragmocone.
Jeletzky [1] characterized phragmoteuthids as having a large tripartite, fanlike pro-ostracum forming the longest portion of the shell, attached to about three-quarters of the circumference of a comparatively small breviconic phragmocone with short camerae and superficially belemnitid-like siphuncle, an absent or much reduced rostrum at the apical part of the phragmocone, belemnite-like arm hooks, an ink sack, beaks resembling those of Recent teuthids, and a muscular mantle.
Donovan (2006), gives a similar description for Phragmoteuthis : Phragmocones as having an apical angle of between 20 and 30 degrees, and relatively few chambers compared with belemnoids; a multi-layered conotheca, thick-walled siphuncle, and a long, three-lobed pro-ostracum as in the Triassic species. Arms are short and bear pairs of slightly curved hooks. [2]
Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids than they are to shelled nautiloids such as the living Nautilus species. The earliest ammonites appeared during the Devonian, with the last species vanishing shortly after the Cretaceous–Paleogene extinction event, during the Danian epoch of the Paleocene.
Subclass Coleoidea, or Dibranchiata, is the grouping of cephalopods containing all the various taxa popularly thought of as "soft-bodied" or "shell-less". Unlike its extant sister group, Nautiloidea, whose members have a rigid outer shell for protection, the coleoids have at most an internal cuttlebone, gladius, or shell that is used for buoyancy or support. Some species have lost their cuttlebone altogether, while in some it has been replaced by a chitinous support structure. A unique trait of the group is the ability to edit their own RNA.
Belemnoids are an extinct group of marine cephalopod, very similar in many ways to the modern squid and closely related to the modern cuttlefish. Like them, the belemnoids possessed an ink sac, but, unlike the squid, they possessed ten arms of roughly equal length, and no tentacles. The name "belemnoid" comes from the Greek word βέλεμνον, belemnon meaning "a dart or arrow" and the Greek word είδος, eidos meaning "form".
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day.
The phragmocone is the chambered portion of the shell of a cephalopod. It is divided by septa into camerae.
Orthocerida is an order of extinct Orthoceratoid cephalopods also known as the Michelinocerida that lived from the Early Ordovician possibly to the Late Triassic. A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous. They were most common however from the Ordovician to the Devonian.
The Bactritida are a small order of more or less straight-shelled (orthoconic) cephalopods that first appeared during the Emsian stage of the Devonian period with questionable origins in Pragian stage before 409 million years ago, and persisted until Carnian pluvial event in the upper middle Carnian stage of the Triassic period. They are considered ancestors of the ammonoids, as well as of the coleoids.
Boletzkyida is a primitive order of teuthid coleoid cephalopod: the boletzkyids are thought to be the earliest forms of coleoid cephalopods, and appear to form a link between nautiloid orthocerids and more advanced coleoids. Boletzkyida was named and described by Bandel, Reitner, and Sturmer in 1983 (B.R.&S) from specimens found in the Lower Devonian black slate in Germany.
Actinoceras is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician. It is an extinct genus of nautiloid cephalopod that thrived in the warm waters of the United States during the Paleozoic era.
Belemnitella is a genus of belemnite from the Late Cretaceous of Europe and North America. Belemnitella was a squidlike animal, probably related to the ancestors of modern squids and cuttlefish. The shell was internal. The rostrum or guard is found the most often and possesses a distinctive slit at its ventral surface and a ridge on the dorsal surface. The phragmocone has a small protoconch at its tip, and fit into the cavity and the guard. It also had septa and a ventral siphuncle within the phragmocone, projecting forward as a beak-like blade.
Belemnotheutis is an extinct coleoid cephalopod genus from the middle and upper Jurassic, related to but morphologically distinct from belemnites. Belemnotheutis fossils are some of the best preserved among coleoids. Remains of soft tissue are well-documented in some specimens, even down to microscopic muscle tissue. In 2008, a group of paleontologists even recovered viable ink from ink sacs found in several specimens.
Phragmoteuthis is a genus of extinct coleoid cephalopod known from the late Triassic to the lower Jurassic. Its soft tissue has been preserved; some specimens contain intact ink sacs, and others, gills. It had an internal phragmocone and unknown numbers of arms.
Acanthoteuthis is a belemnite genus, a squid-like cephalopod with an internal shell from the Late Jurassic Epoch, related to modern coleoids.
Kobyashiceras is an extinct genus in the cephalopod order Actinocerida, from Lower Devonian marine sediments in Japan. The type and sole included species is Kobayashiceras gifuense. The generic name honors the late Dr. Teiichi Kobayashi who contributed greatly to the study of Paleozoic cephalopods. The specific name is derived from Gifu, the name of the prefecture in which the type locality is found.
The cephalopods have a long geological history, with the first nautiloids found in late Cambrian strata, and purported stem-group representatives present in the earliest Cambrian lagerstätten.
Hematitida is a group of coleoid cephalopods known from the early Carboniferous Period. They are the oldest definite coleoids, although there are controversial claims for even older coleoids from the Devonian. Fossil hematitidans have so far been found only in Arkansas and Utah of the United States. The only family described so far is Hematitidae.
Belemnitida is an extinct order of squid-like cephalopods that existed from the Late Triassic to Late Cretaceous. Unlike squid, belemnites had an internal skeleton that made up the cone. The parts are, from the arms-most to the tip: the tongue-shaped pro-ostracum, the conical phragmocone, and the pointy guard. The calcitic guard is the most common belemnite remain. Belemnites, in life, are thought to have had 10 hooked arms and a pair of fins on the guard. The chitinous hooks were usually no bigger than 5 mm (0.20 in), though a belemnite could have had between 100 and 800 hooks in total, using them to stab and hold onto prey.
Reedsoceras is a genus of large discosorids (Nautiloidea) in the family Westonoceratidae from the middle and upper Ordovician of North America.
Simardoceras is a genus in the discosorid family Westonoceratidae from the Middle Ordovician of Quebec.
Clarkeiteuthis is a genus of extinct belemnoid cephalopod known from the lower Jurassic in Germany and England. Described two species, C. conocauda and C. montefiorei are originally described as species of phragmoteuthid Phragmoteuthis, but got their own genus and moved to Diplobelida.
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