Lituitida Temporal range: | |
---|---|
Trilacinoceras | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Subclass: | † Orthoceratoidea |
Order: | † Lituitida |
Lituitida is an order of orthoceratoid cephalopods. They correspond to the family Lituitidae of the Treatise (Furnish & Glenister, 1964), reranked as an order and combined with other orthoceratoids. [1] They are considered to be more closely related to the Orthocerida than to the Ascocerida or Pseudorthocerida which are also included.
Lituitids are characterized by smooth to annulate shells in which the juvenile portion near the apex is coiled or cyrtoconic. The adult portion is straight or slightly sigmoidal and may be expanded. Growth lines show a deep hyponomic sinus and lateral ocular sinuses. The siphuncle is subcentral with layered connecting rings. Cameral deposits are prevalent and concentrated more ventrally. [2] The apex is small and spherical and lacks a cicatrix. [1] The Lituitida, which were originally included in the Tarphycerida as the Lituitidae (Furnish and Glenister) have their beginnings in the Early Ordovician with forms like Ancistroceras . Their diversity increased after the late Arenigian extinction (Floian-Dapingian) at the end of the Canadian Epoch and take a tumble following the early Cincinnatian (Sandbian) in the early Late Ordovician, from which the group never recovers. [3]
It is thought that they had a planktonic juvenile stage in their life cycle and that mature adults lived in the shallow waters of the open ocean. [3]
Some 8 genera have been described, all from the Baltic region of northern Europe. One genus, Holmiceras, is limited to the Lower Ordovician and one, Ancistroceras, comes from both the lower and middle of the period. Five genera included the type genus, Lituites , are known only from the Middle Ordovician and one, Tyrioceras, is known from the Middle and Upper Ordovician, the last of its kind.
One study argued that Lituitida (also considered a suborder, Lituitina) was closely related to the proposed order Pallioceratida (or suborder Pallioceratina). Under this hypothesis, they would collectively form the superorder Astrovioidea (or order Astroviida), which was diagnosed based on extensive cameral deposits which partially obliterate the connecting rings. [4] [5] However, the validity of Pallioceratida/Pallioceratina or Astroviodea/Astroviida are not universally accepted. [6] [7]
Mutvei, H (2002). "Connecting ring structure and its significance for classification of the orthoceratid cephalopods". Acta Palaeontologica Polonica. 47 (1): 157–168.
Lituites is an extinct nautiloid genus from the Middle Ordovician and type for the Lituitidae that in some more recent taxonomies has been classified with the orthocerids and listed under the order Lituitida. Fossils have been found in New York, Argentina, Norway, Sweden, Estonia, and China.
Endocerida is an extinct nautiloid order, a group of cephalopods from the Lower Paleozoic with cone-like deposits in their siphuncle. Endocerida was a diverse group of cephalopods that lived from the Early Ordovician possibly to the Late Silurian. Their shells were variable in form. Some were straight (orthoconic), others curved (cyrtoconic); some were long (longiconic), others short (breviconic). Some long-shelled forms like Endoceras attained shell lengths close to 6 metres (20 ft). The related Cameroceras is anecdotally reported to have reached lengths approaching 9 metres (30 ft), but these claims are problematic. The overwhelming majority of endocerids and nautiloids in general are much smaller, usually less than a meter long when fully grown.
Orthoceratidae is an extinct family of actively mobile carnivorous cephalopods, subclass Nautiloidea, that lived in what would be North America, Europe, Asia, Africa, and Australia from the Ordovician through Triassic from 490—203.7 mya, existing for approximately 286.4 million years.
Orthocerida, also known as the Michelinocerida, is an order of extinct orthoceratoid cephalopods that lived from the Early Ordovician possibly to the Late Triassic. A fossil found in the Caucasus suggests they may even have survived until the Early Cretaceous, and the Eocene fossil Antarcticeras is sometimes considered a descendant of the orthocerids although this is disputed. They were most common however from the Ordovician to the Devonian.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician to the Middle Devonian. Some, such as Aphetoceras and Estonioceras, are loosely coiled and gyroconic; others, such as Campbelloceras, Tarphyceras, and Trocholites, are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular, as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably Bassleroceras, around the end of the Gasconadian through forms like Aphetoceras. Close coiling developed rather quickly, and both gyroconic and evolute forms are found in the early middle Canadian.
Anthoceras is a genus of straight, annulated, proterocamerioceratid molluscs from the Lower Ordovician, found in North America, North-Western Australia, and Siberia. The cross section is circular, the siphuncle moderately large, and marginal. Segments are constricted ; septal necks hemichoantici to subholochoantic ; connecting rings thick. Endocones are long and slightly asymmetric.
Bactroceras is a genus of orthoceratoid cephalopods that lived during the early Middle Ordovician, from about 472—464 mya, existing for approximately 8 million years.
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.
Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars, extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer.
The Bassleroceratidae is a family of gradually expanding, smooth ellesmerocerids with a slight to moderate exogastric curvature, subcircular to strongly compressed cross section, and ventral orthochaonitc siphuncle. The ventral side is typically more sharply rounded than the dorsal side and septa are close spaced. Connecting rings are thick and slightly expanded into the siphuncle, making the segments slightly concave; characteristic of the Ellesmerocerida.
The Orthocerataceae is a superfamily of orthocerid cephalopods that lived from the late Early Ordovician to the Early Cretaceous, but is no longer in general use.
Ancistroceras is one of the two ancestral lituitids from the late Early Ordovician (Arenigian). The other being Holmiceras.
Holmiceras is one of two ancestral lituitids from the late Early Ordovician (Arenigian). The other being Ancistroceras.
The Lituitidae are a family of evolved tarphycerids characterized by a long orthoconic section that follows a coiled juvenile portion at the apex, along with a generally tubular siphuncle, which like that of the barrandeocerids is composed of thin connecting rings.
Plectronoceratoidea is a superorder or subclass containing primitive nautiloids from the Late Cambrian and Early Ordovician. This group is best considered a paraphyletic grade of early cephalopods, as it contains the ancestors of subsequent post-Cambrian cephalopod orders.
Ophioceras is a genus of closely coiled tarphycerid nautiloid cephalopods, the sole representatives of the family Ophidioceratidae, characterized by an evolute shell with narrow, subrounded, annulated whorls and a subcentral siphuncle composed of thin connecting rings that show no evidence of layering. The mature body chamber is strongly divergent and is the longest proportionally of any tarphycerid. The aperture has a deep hyponomic sinus and ocular sinuses, and so resembles some lituitids.
Cyrtocerinida is an order of Ordovician nautiloid cephalopods. The order includes the families Cyrtocerinidae and Eothinoceratidae, as well as the genera Bathmoceras and Rummoceras.
Multiceratoidea is a major subclass or superorder of Paleozoic nautiloid cephalopods. Members of this group can be characterized by nautilosiphonate connecting rings, with an organic inner layer and outer layer of calcitic spherules and blades, similar to the modern nautilus. The earliest-diverging multiceratoids have oncomyarian muscle scars, though several orders trend towards a ventromyarian condition. Multiceratoid shells are generally short and curled, with a relatively small aperture (opening). Cameral deposits are never found among the multiceratoids, though several orders are known to bear endosiphuncular deposits within their siphuncles.
Riocerida, originally named as Rioceratida, is an extinct order of Ordovician nautiloid cephalopods. They were the earliest-diverging members of the major group Orthoceratoidea, with which they share dorsomyarian muscle scars. However, they also possess several plesiomorphic ("primitive") traits which are absent in other orthoceratoids: Cameral and endosiphuncular deposits are absent, and the siphuncle has a marginal position along the lower edge of the shell. The conch shape is slender, usually orthoconic (straight) or weakly cyrtoconic. The monophyly of Riocerida as a unique group is plausible but difficult to confirm, and they may instead be a paraphyletic grade or polyphyletic assemblage ancestral to all later orthoceratoids.