Centipedes are predatory arthropods belonging to the class Chilopoda of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful stings, injecting their venom through pincer-like appendages known as forcipules or toxicognaths, which are actually modified legs instead of fangs. Despite the name, no centipede has exactly 100 pairs of legs; number of legs ranges from 15 pairs to 191 pairs, always an odd number.
Pachymerium is a genus of centipedes in the family Geophilidae. Centipedes in this genus feature elongate heads, sternal pores in a pair of anterior groups and a posterior transverse band, and scattered coxal pores; the forcipular coxosternite is broad, and the ultimate article of the forcipule has a conspicuous basal denticle. These centipedes range from about 2 cm to about 8 cm in length and have 37 to 79 pairs of legs. The Chilean species Pachymerium armatum measures only 20 mm in length and has only 37 leg pairs, the minimum number observed in this genus. The Russian species P. minutum is also notable for its small size and its modest number of legs. The Portuguese species P. coiffaiti is notable for its large size and many legs. Females of the large Turkish species P. serratum can reach 78 mm in length and have as many as 79 leg pairs, the maximum number observed in this genus. This genus contains the following species:
The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea containing the mostly defunct clades Aphilodontidae, Dignathodontidae, Linotaeniidae, Chilenophilinae, and Macronicophilidae.
Geophilus is a large, heterogeneous genus of soil centipedes in the family Geophilidae largely considered to be synonymous with Brachygeophilus. This genus is a mostly holarctic and characterized by a claw-shaped ultimate pretarsus, anterior porefields, complete or nearly complete coxo-pleural sutures at the prosternum, and incomplete chitin-lines. Centipedes in this genus feature slightly elongate heads and labral intermediate parts with tubercles, the forcipules are usually poorly elongate with a single small tubercle at the base of each tarsungulum, and the anterior trunk metasternites usually have an anterior medial socket and a posterior transversally elongate pore-field. The generic name first appeared in Brewster's Edinburgh Encyclopaedia in 1814 as Geophilus electricus.
Linotaeniidae are a monophyletic clade of soil centipedes in the family Geophilidae found mostly in the temperate regions of the Holarctic as well as the south Andes. Species in the clade Linotaeniidae are characterized by a body that usually tapers toward the anterior tip; mandibles with a single pectinate lamella; second maxillae with coxo-sternite usually undivided and claws without projections; forcipular segment short, with tergite remarkably wide, forcipules evidently tapering; coxal organs opening through distinct pores on the ventral surface of the coxo-pleura. The number of legs in this clade varies within as well as among species and ranges from as few as 31 pairs of legs to as many as 83 leg pairs. Compared to most families in the suborder Adesmata, this clade features a modest number of leg-bearing segments and limited variation in this number within each species.
Schendylidae is a family of centipedes in the order Geophilomorpha.
Chileana is a genus of soil centipedes in the clade Linotaeniidae and family Geophilidae found in southern Chile. It currently has only one species, C. araucanensis. Females of this species are about 30mm long, with a pale yellow body and a red head; bearing 12–15 pleural pores; long, tapering antennae with sparse basal sections and rather hairy distal sections; and a labrum with four median tubercles bearing a few cilia on the sides. Males have 10 pleural pores, thick ultimate legs armed with claws, and 43 leg pairs.
Ballophilidae is a monophyletic group of centipedes belonging to the order Geophilomorpha and superfamily Himantarioidea. Authorities now dismiss this group as a family, citing phylogenetic analysis, and instead refer to this clade as Ballophilinae, a possible subfamily within the family Schendylidae. Centipedes in this clade differ from others in the family Schendylidae by having bodies tapered toward the anterior tip, short heads, slender forcipules that are well apart, and an undivided lamina for the female gonopods.
Ballophilus is the largest genus of centipedes in the clade Ballophilidae. Species in this genus are found in tropical and subtropical regions, mainly in Africa and southeast Asia.
Ballophilus differens is a species of centipede in the genus Ballophilus. It is found in Angola. The original description of this species is based on specimens reaching up to 15 mm in length with 47 pairs if legs.
Caritohallex is a monotypic genus of centipede with only one species, Caritohallex minyrrhopus, found in Panama and the Lesser Antilles. This species has slender antennae, no coxal pores, and a single tarsal article on the ultimate legs. The original description of this species is based on a male holotype measuring about 10 mm in length with 39 pairs of legs and a female paratype measuring 8 mm in length with 43 leg pairs.
Geophilus virginiensis is a species of soil centipede in the family Geophilidae found in Virginia. It grows up to 50 millimeters in length, has 49–57 leg pairs, and is tawny yellow in color.
Geophilus richardi is a species of soil centipede in the family Geophilidae found in France, Italy, and the Ionian Islands. Females of this species have 33 pairs of legs; males have 29 or 31. This species is one of only two in the family Geophilidae to include centipedes with as few as 29 leg pairs. This species grows up to 10 millimeters long, has no carpophagus pit or pore-fields, and has a gradually tapering, curved pretarsus of the second maxillae. G. richardi lacks typical ventral pores between 2–4mm. The sternites instead bear a small number of pores between 0.5–1mm that differ from micropores, which are unbounded by a cuticular ring. These are possibly the remnants of typical ventral pores, their smaller size being a byproduct of overall miniaturization.
Sepedonophilus is a genus of three species of centipedes in the family Geophilidae. This genus is endemic to Australia, and was first described by Austrian myriapodologist Carl Attems in 1909. Centipedes in this genus feature scattered coxal pores and have no sternal pores; the lateral parts of the labrum almost touch medially, and the intermediate part is inconspicuous; the coxosternite of the second maxillae have peculiar anterior projections. These centipedes range from about 2 cm to 5 cm in length and have 49 to 79 pairs of legs. The smallest species in this genus, Sepedonophilus hodites, measures only 18 mm in length and has only 49 leg pairs, the minimum number recorded in this genus, whereas the largest species, S. perforatus, measures 50 mm in length and has 79 leg pairs, the maximum number recorded in this genus.
Pachymerinus is a genus of centipedes in the family Geophilidae. It was described by Italian entomologist Filippo Silvestri in 1905. Centipedes in this genus feature elongate heads, elongate forcipules with denticles, scattered coxal pores, and few sternal pores, if any; the intermediate part of the labrum is narrow and has no denticles. These centipedes range from about 3 cm to 8 cm in length, have 47 to 81 pairs of legs, and are found in Chile and southeast Australia. The Australian species Pachymerinus froggatti is notable for its relatively small size, measuring only 28 mm in length. The larger Chilean species P. porteri measures 46 mm in length but can have as few as 47 leg pairs, the minimum number recorded in this genus. The Chilean species P. pluripes measures only 32 mm in length but can have 79 or 81 leg pairs, the maximum number recorded in this genus. The Chilean species P. canaliculatus is known from a female specimen with 75 leg pairs and is notable for its large size, measuring 75 mm in length.
Queenslandophilus is a genus of centipedes in the family Geophilidae. It was described by German myriapodologist Karl Wilhelm Verhoeff in 1925. Centipedes in this genus feature second maxillae with sclerotized ridges on the coxosternite, no sternal pores, and coxopleura many scattered pores; the lateral parts of the labrum almost touch medially, and the ultimate legs most often have claws. These centipedes range from 1 cm to 6 cm in length, have 37 to 75 pairs of legs, and are found in Australia, Japan, and North America. The Japanese species Queenslandophilus monoporus and Q. macropalpus are notable for their small sizes, measuring only 10 mm and 15 mm in length, respectively. The species Q. elongatus, found in California, is notable for its large size, ranging from 40 mm to 63 mm in length. The species Q. macropalpus has only 37 leg pairs, the minimum number recorded in this genus, whereas Q. elongatus has 73 to 75 leg pairs, the maximum number recorded in this genus.
Schendyla is a genus of centipedes in the family Schendylidae. It was described by Danish entomologists Vilhelm Bergsøe and Frederik Vilhelm August Meinert in 1866. Centipedes in this genus feature only a few spines on the claws of the second maxillae, two pores on each coxopleuron, and ultimate legs without claws. These centipedes range from 1 cm to about 4 cm in length and have from 31 to 57 pairs of legs. The species Schendyla verneri is notable for its small size and for having only 31 leg pairs, the minimum recorded in this genus. Other small species with notably few legs in this genus include S. walachica, S. dalmatica, S. dentata, S. gracillima. and S. armata. The species S. monoeci measures only 17.5 mm in length but can have from 51 to as many as 57 leg pairs, the maximum number found in this genus. Descriptions of the species S. vizzavonae report no more than 51 leg pairs, but this species is notable for its relatively large size, reaching 45 mm in length.
Plesioschendyla is a monotypic genus of centipedes in the family Schendylidae. It is endemic to New Caledonia, a French overseas territory in Melanesia. Its sole species is Plesioschendyla confossa. It was described by French entomologist Henri Ribaut in 1923.
Pectiniunguis is a genus of centipedes in the family Schendylidae. It was described by American naturalist Charles Harvey Bollman in 1889. Centipedes in this genus feature second maxillae with claws fringed by two rows of filaments, transversally elliptical sternal pore-fields on almost all trunk segments, two pores on each coxopleuron, and ultimate legs without claws. These centipedes range from 16 mm to 67 mm in length, have 35 to 73 pairs of legs, and are found in the tropical and subtropical regions of the Americas, Fiji, and west Africa. The African species Pectiniunguis minutus is notable not only for being the smallest in this genus but also for having as few as 35 leg pairs, the minimum number in this genus. The Brazilian species P. ducalis is notable not only for being the largest in this genus but also for having as many as 73 leg pairs, the maximum number in this genus.
Marsikomerus is a genus of centipedes in the family Schendylidae. It was described by Austrian myriapodologist Carl Attems in 1938. Species in this genus are found in Mexico and in the United States.
