Caladenia attingens subsp. effusa

Granite mantis orchid
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Monocots
Order:	Asparagales
Family:	Orchidaceae
Subfamily:	Orchidoideae
Tribe:	Diurideae
Genus:	Caladenia
Species:	C. attingens
Subspecies:	C. a. subsp. effusa
Trinomial name
	Caladenia attingens subsp. effusa; A.P.Br. & G.Brockman [1]

Last updated November 03, 2019

Caladenia attingens subsp. effusa, commonly known as granite mantis orchid, is a subspecies of orchid endemic to the south-west of Western Australia. It has a single erect, hairy leaf and one or two green, yellow and red flowers. It differs from the other subspecies of Caladenia attingens in having smaller flowers with lateral sepals which are not upswept and a labellum which is less than 15 mm (0.6 in) wide.

Endemism is the ecological state of a species being unique to a defined geographic location, such as an island, nation, country or other defined zone, or habitat type; organisms that are indigenous to a place are not endemic to it if they are also found elsewhere. The extreme opposite of endemism is cosmopolitan distribution. An alternative term for a species that is endemic is precinctive, which applies to species that are restricted to a defined geographical area.

Western Australia is a state occupying the entire western third of Australia. It is bounded by the Indian Ocean to the north and west, and the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Australia is Australia's largest state, with a total land area of 2,529,875 square kilometres, and the second-largest country subdivision in the world, surpassed only by Russia's Sakha Republic. The state has about 2.6 million inhabitants – around 11 percent of the national total – of whom the vast majority live in the south-west corner, 79 per cent of the population living in the Perth area, leaving the remainder of the state sparsely populated.

A sepal is a part of the flower of angiosperms. Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom. The term sepalum was coined by Noël Martin Joseph de Necker in 1790, and derived from the Greek σκέπη, a covering.

Description

Caladenia attingens subsp. effusa is a terrestrial, perennial, deciduous, herb with an underground tuber and a single hairy leaf, 50–200 mm (2–8 in) long and 5–11 mm (0.2–0.4 in) wide. One or two green, yellow and red flowers 40–60 mm (2–2 in) long and 30–40 mm (1–2 in) wide are borne on a stalk 120–250 mm (5–10 in) tall. The sepals and petals have thin, brown, club-like glandular tips 10–12 mm (0.4–0.5 in) long. The dorsal sepal is erect, 30–40 mm (1–2 in) long and about 2 mm (0.08 in) wide and the lateral sepals are 30–40 mm (1–2 in) long, 1.5–2.5 mm (0.06–0.1 in) wide, downswept near the base then curving forwards and slightly upwards. The petals are 20–30 mm (0.8–1 in) long, 1–2 mm (0.04–0.08 in) wide and downswept. The labellum is 9–15 mm (0.4–0.6 in) long, 12–15 mm (0.5–0.6 in) wide and green with a dark red tip. The sides of the labellum have narrow teeth and there are four or more rows of crowded, red calli along its centre including near its tip. Flowering occurs from August to early October. This subspecies differs from the other two subspecies in having lateral sepals that are not strongly upswept and a labellum that is less than 15 mm (0.6 in) wide.^[2]^[3]^[4]

A terrestrial plant is a plant that grows on, in, or from land. Other types of plants are aquatic, epiphytic and lithophytic.

A perennial plant or simply perennial is a plant that lives more than two years. Some sources cite perennial plants being plants that live more than three years. The term is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widely used to distinguish plants with little or no woody growth from trees and shrubs, which are also technically perennials.

In the fields of horticulture and botany, the term deciduous means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, after flowering; and to the shedding of ripe fruit.

Taxonomy and naming

Caladenia attingens was first formally described in 2001 by Stephen Hopper and Andrew Phillip Brown and the description, including of two subspecies, was published in Nuytsia .^[5] In 2015, Andrew Brown and Garry Brockman described subspecies effusa which, before formal description, was known as Caladenia attingens subsp. 'granite'. The description of the new subspecies was published in a later edition of Nuytsia.^[1] The name effusa refers to the lateral sepals which spread outwards rather than being sickle-shaped as in the other two subspecies.^[4]

Stephen Donald Hopper AC FLS FTSE is a Western Australian botanist, graduated in Biology, specialising in conservation biology and vascular plants. He has written eight books, and has over 200 publications to his name. He was Director of Kings Park in Perth for seven years, and CEO of the Botanic Gardens and Parks Authority for five. He is currently Foundation Professor of Plant Conservation Biology at The University of Western Australia. He was Director of the Royal Botanic Gardens, Kew from 2006 to 2012.

Andrew Phillip Brown is a conservation biologist and taxonomist at the Western Australian Department of Environment and Conservation. He is also curator of Orchidaceae and Myoporaceae at the Western Australian Herbarium and a foundation member of the Australian Orchid Foundation and the Western Australia Native Orchid Study and Conservation Group. He is the author of more than 100 journal articles and seven books on the flora of Western Australia including a field guide to the eremophilas of that state.

Nuytsia is a peer-reviewed scientific journal published by the Western Australian Herbarium. It publishes papers on systematic botany, giving preference to papers related to the flora of Western Australia. Nearly twenty percent of Western Australia's plant taxa have been published in Nuytsia. The journal was established in 1970 and has appeared irregularly since. The editor-in-chief is Kevin Thiele.

Distribution and habitat

The granite mantis spider orchid is found between Peak Charles and Balladonia in the Mallee biogeographic region where it grows in shallow soil on granite outcrops.^[2]^[3]^[6]

Balladonia is a small roadhouse community located on the Eyre Highway in Western Australia. It is the first stop east of Norseman on the long journey east across the Nullarbor Plain. Between Balladonia and Caiguna is a 146.6-kilometre (91.1 mi) stretch of the highway which is one of the longest straight stretches of road in the world.

Mallee, also known as Roe Botanical District, is a biogeographic region in southern Western Australia. Located between the Esperance Plains, Avon Wheatbelt and Coolgardie regions, it has a low, gently undulating topography, a semi-arid mediterranean climate, and extensive Eucalyptus mallee vegetation. About half of the region has been cleared for intensive agriculture. Recognised as a region under the Interim Biogeographic Regionalisation for Australia (IBRA), it was first defined by John Stanley Beard in 1980.

Granite A common type of intrusive, felsic, igneous rock with granular structure

Granite is a common type of felsic intrusive igneous rock that is granular and phaneritic in texture. Granites can be predominantly white, pink, or gray in color, depending on their mineralogy. The word "granite" comes from the Latin granum, a grain, in reference to the coarse-grained structure of such a holocrystalline rock. Strictly speaking, granite is an igneous rock with between 20% and 60% quartz by volume, and at least 35% of the total feldspar consisting of alkali feldspar, although commonly the term "granite" is used to refer to a wider range of coarse-grained igneous rocks containing quartz and feldspar.

Conservation

Caladenia attingens subsp. effusa is classified as "not threatened" by the Western Australian Government Department of Parks and Wildlife.^[6]

The Department of Parks and Wildlife (DPaW) was the department of the Government of Western Australia responsible for managing lands described in the Conservation and Land Management Act 1984 and implementing the state's conservation and environment legislation and regulations. The minister responsible for the department was the Minister for the Environment.

Related Research Articles

Caladenia longicauda is a species of plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It is distinguished by its large leaf and by its up to five large, white flowers which have drooping sepals and petals with long, thickish brown "tails".

Caladenia longicauda subspecies borealis, commonly known as daddy-long-legs spider orchid is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to three mostly white flowers with long, drooping sepals and petals.

Caladenia attingens, commonly known as mantis orchids, are plants in the orchid family Orchidaceae and are endemic to the south-west of Western Australia. There are three subspecies, each of which has a single hairy leaf and one or two brightly coloured flowers with upswept sepals and a labellum with long, comb-like fringes. The subspecies differ in size, distribution and habitat.

Caladenia bicalliata subsp. cleistogama, commonly known as the shy limestone spider orchid or sandhill spider orchid, is a plant in the orchid family Orchidaceae and is native to the south-west of Western Australia. It has a single erect, hairy leaf and one or two cream-coloured flowers which are smaller than those in subspecies bacalliata.

Caladenia corynephora, commonly known as the club-lipped spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single erect, hairy leaf and one or two greenish-yellow and red flowers which have a labellum with a club-like tip. It is the only Western Australian caladenia with a clubbed labellum.

Caladenia denticulata subsp. denticulata, commonly known as the yellow spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single erect, hairy leaf and one or two yellowish flowers which have a white labellum with pale red markings.

Caladenia denticulata subsp. albicans, commonly known as the alabaster spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single erect, hairy leaf and one or two pale creamy-white flowers which have a white labellum with red markings. The only known population occurs in a small area near Arrowsmith.

Caladenia denticulata subsp. rubella, commonly known as the clumped spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single erect, hairy leaf and one or two dull or pinkish red flowers with prominent dull red markings and with a white labellum with red markings.

Caladenia flava subsp. sylvestris, commonly known as the karri cowslip orchid, is a species of orchid endemic to the south-west of Western Australia. It has a single, hairy leaf and up to three pale yellow and cream-coloured flowers which are white near the tips of the sepals and petals and marked with bright red or pink.

Caladenia longicauda subsp. crassa, commonly known as the Esperance white spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to three mostly white flowers with long, broad, spreading lateral sepals and petals, a relatively broad labellum with short, narrow teeth. It grows on the south coast between Bremer Bay and the Cape Arid National Park.

Caladenia longicauda subsp. eminens, commonly known as the stark white spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to three large, bright white flowers with long, broad, spreading lateral sepals and petals, a relatively broad labellum with short, narrow teeth. It is a relatively common orchid found in a broad band, mainly between Tenterden and Jerramungup.

Caladenia longicauda subsp. longicauda, commonly known as the white spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to four, mostly white flowers with long drooping, thread-like ends on the sepals and petals. It grows in woodland and forest.

Caladenia applanata subsp. applanata, commonly known as the broad-lipped spider orchid, is a species of orchid endemic to the south-west of Western Australia. It is a relatively common orchid with a single erect, hairy leaf and up to three flowers which may be red, cream, green or yellow and have a broad, flattened, red-tipped labellum.

Caladenia attingens subsp. attingens, commonly known as the forest mantis orchid or sneezing spider orchid, is a species of orchid endemic to the south-west of Western Australia. It is a relatively common orchid with a single erect, hairy leaf and one or two green, yellow and red flowers. It is similar to the fringed mantis orchid but has smaller flowers and has a more southerly distribution.

Caladenia attingens subsp. gracillima, commonly known as the small mantis orchid, is a species of orchid endemic to the south-west of Western Australia. It is a relatively common orchid with a single erect, hairy leaf and one or two green, yellow and red flowers. It differs from subspecies attingens in having smaller flowers and a more easterly distribution.

Caladenia longicauda subsp. merrittii, commonly known as Merritt's white spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to three large, mainly white flowers with very long, drooping lateral sepals and petals, and a white broad labellum with relatively short labellum teeth. It is one of the largest spider orchids.

Caladenia longicauda subsp. redacta, commonly known as the tangled white spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to three large, mainly white flowers with long, drooping lateral sepals and petals. It is most similar to subspecies eminems but has smaller flowers and shorter teeth on the side of the labellum.

Caladenia longicauda subsp. extrema, commonly known as the late white spider orchid or Seaton Ross spider orchid is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and one or two mainly white flowers with long, mostly spreading lateral sepals and petals. It is a relatively rare orchid which is similar to the tangled white spider orchid but has larger flowers and a later flowering period.

Caladenia longicauda subsp. insularis, commonly known as the island white spider orchid is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to four yellowish-white flowers with long, mostly spreading lateral sepals and petals. It is a relatively rare, self-pollinating subspecies and often flowers which are in bud, open and finished are seen on a single plant.

Caladenia longicauda subsp. minima, commonly known as the little white spider orchid, is a plant in the orchid family Orchidaceae and is endemic to the south-west of Western Australia. It has a single hairy leaf and up to three mainly white flowers with long, drooping lateral sepals and petals. It is a relatively rare orchid which is similar to the daddy-long-legs spider orchid but has smaller flowers and a more easterly distribution.

References

1 2 "Caladenia attingens subsp. effusa". APNI. Retrieved 17 February 2017.
1 2 Brown, Andrew; Dixon, Kingsley; French, Christopher; Brockman, Garry (2013). Field guide to the orchids of Western Australia : the definitive guide to the native orchids of Western Australia. Simon Nevill Publications. p. 119. ISBN 9780980348149.
1 2 Hoffman, Noel; Brown, Andrew (2011). Orchids of South-West Australia (3rd ed.). Gooseberry Hill: Noel Hoffman. p. 146. ISBN 9780646562322.
1 2 Brown, Andrew Phillip; Brockman, Garry (2015). "New taxa of Caladenia (Orchidaceae) from south-west Western Australia". Nuytsia. 25: 61–65.
↑ "Caladenia attingens". APNI. Retrieved 17 February 2017.
1 2 "Caladenia attingens subsp. effusa". FloraBase . Western Australian Government Department of Parks and Wildlife.

This page is based on this Wikipedia article
Text is available under the CC BY-SA 4.0 license; additional terms may apply.
Images, videos and audio are available under their respective licenses.

[APNI-1] 1 2 "Caladenia attingens subsp. effusa". APNI. Retrieved 17 February 2017.

[Brown-2] 1 2 Brown, Andrew; Dixon, Kingsley; French, Christopher; Brockman, Garry (2013). Field guide to the orchids of Western Australia : the definitive guide to the native orchids of Western Australia. Simon Nevill Publications. p. 119. ISBN 9780980348149.

[Hoffman-3] 1 2 Hoffman, Noel; Brown, Andrew (2011). Orchids of South-West Australia (3rd ed.). Gooseberry Hill: Noel Hoffman. p. 146. ISBN 9780646562322.

[Nuytsia(2)-4] 1 2 Brown, Andrew Phillip; Brockman, Garry (2015). "New taxa of Caladenia (Orchidaceae) from south-west Western Australia". Nuytsia. 25: 61–65.

[APNI(1)-5] "Caladenia attingens". APNI. Retrieved 17 February 2017.

[FloraBase-6] 1 2 "Caladenia attingens subsp. effusa". FloraBase . Western Australian Government Department of Parks and Wildlife.