Ceratotherium neumayri | |
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Skull | |
Scientific classification ![]() | |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Family: | Rhinocerotidae |
Genus: | Ceratotherium |
Species: | †C. neumayri |
Binomial name | |
†Ceratotherium neumayri (Osborn, 1900) | |
Synonyms | |
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Ceratotherium neumayri is an extinct species of rhinoceros from the Late Miocene epoch (specifically the Vallesian and Turolian European land mammal ages) of the Balkans (including Greece [1] and Bulgaria [2] ) and Western Asia (including Iran and Anatolia in Turkey). [3]
The species was originally named Atelodus neumayri by Henry Fairfield Osborn in 1900. [4] It is considered part of the tribe Dicerotini (also spelled Diceroti) or subtribe Dicerotina, indicating a close relationship to the extant African rhinoceroses, the black rhinoceros (Diceros bicornis) and the white rhinoceros (Ceratotherium simum). [5]
The generic assignment of neumayri is disputed, having been placed in both Ceratotherium and Diceros by various authors. [6] Some paleontologists have proposed it as a common ancestor to both Ceratotherium and Diceros, [7] [8] while others suggest it represents an early, distinct evolutionary branch not directly ancestral to the modern African rhinos. [5] A 2022 study placed the species in the separate monotypic genus Miodiceros. [5]
The species was a large sized rhinceros, and had two horns, a nasal and a frontal horn. The nasal septum was not ossified. [5]
Analysis of dental microwear patterns on the teeth of C. neumayri suggests that it was a mixed feeder. This diet means it was adaptable, capable of both grazing on grasses and browsing on leaves and twigs from shrubs and trees. [9]
Fossils attributed to C. neumayri have been found across southeastern Europe and Western Asia. Fossils of the species have been found in the Balkans, including Bulgaria [2] and Greece (such as Pikermi, Samos, and Axios Valley). [1] [5] Discoveries also extend into Anatolia and northern Iran, with occurrences in the southern Caucasus. [5]
In 2012, a well-preserved skull from Gülşehir, dating to around 9.2 million years ago was found. This individual is believed to have died due to extreme temperatures from a pyroclastic flow (ignimbrite) associated with a volcanic eruption. [3]
Some authors have suggested that the species was also present in Africa, based on Late Miocene remains found in Tunisia originally attributed to C. douariense. [10]