Empis snoddyi | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Diptera |
Family: | Empididae |
Genus: | Empis |
Species: | E. snoddyi |
Binomial name | |
Empis snoddyi Steyskal, 1969 | |
Empis snoddyi is a species of dance flies in the family Empididae. [1] [2] [3] [4] It is present in the United States, specifically mountainous areas from southern Virginia to northern Georgia. [5] These flies are known for their distinctive mating ritual in which the males offer the females a large empty sack filled with hundreds of silk bubbles. [6]
The thorax and head are darkly colored with grey frosted areas. Eyes are nearly holoptic, meaning that they almost meet in the middle of the head, but there is a slight separation. Legs are yellow with a dark ring where the femur ends. Wings are semi-clear with brown veins throughout. Males and females look quite similar. Two differences are that the separation between the eyes is usually wider in males, and the bristles on the upper part of the leg are shorter in males. [5]
Empis snoddyi flies have evolved a mating ritual in which males provide an empty sack filled with silk bubbles as nuptial gifts to females. Other species of Empididae provide nuptial gifts in the form of prey, wrapped prey, or sacks filled with prey, so it is likely that the empty balloons provided by Empis snoddyi relic related to these nutritious nuptial gifts. It has been hypothesized that these balloons are a representation of the male’s fitness, and it has been shown that large males with intermediately sized sacks have the highest mating rate. [6]
The prothoracic basitarsus (a leg segment) in the male produces the silk for the balloon. [7] The balloon complex is made up of air-filled polygons made of silk. The males are able to vary their output of silk, which leads an irregular pattern of different sized polygons. [8]
A spermatophore or sperm ampulla is a capsule or mass containing spermatozoa created by males of various animal species, especially salamanders and arthropods, and transferred in entirety to the female's ovipore during reproduction. Spermatophores may additionally contain nourishment for the female, in which case it is called a nuptial gift, as in the instance of bush crickets. In the case of the toxic moth Utetheisa ornatrix, the spermatophore includes sperm, nutrients, and pyrrolizidine alkaloids which prevent predation because it is poisonous to most organisms. However, in some species such as the Edith's checkerspot butterfly, the "gift" provides little nutrient value. The weight of the spermatophore transferred at mating has little effect on female reproductive output.
Zygiella x-notata, sometimes known as the missing sector orb weaver or the silver-sided sector spider, is a spider species in the family Araneidae. They are solitary spiders, residing in daily-spun orb webs. Z. x-notata is a member of the genus Zygiella, the orb-weaving spiders. The adult female is easily recognized by the characteristic leaf-like mark on her posterior opisthosoma, caudal to the yellow-brown cephalothorax.
Dolichopodidae, the long-legged flies, are a large, cosmopolitan family of true flies with more than 7,000 described species in about 230 genera. The genus Dolichopus is the most speciose, with some 600 species.
Spider behavior refers to the range of behaviors and activities performed by spiders. Spiders are air-breathing arthropods that have eight legs and chelicerae with fangs that inject venom. They are the largest order of arachnids and rank seventh in total species diversity among all other groups of organisms which is reflected in their large diversity of behavior.
Empididae is a family of flies with over 3,000 described species occurring worldwide in all the biogeographic realms but the majority are found in the Holarctic. They are mainly predatory flies like most of their relatives in the Empidoidea, and exhibit a wide range of forms but are generally small to medium-sized, non-metallic and rather bristly.
Larinioides sclopetarius, commonly called bridge-spider or gray cross-spider, is a relatively large orb-weaver spider with Holarctic distribution. These spiders are located in Europe and have been observed as south as the Mediterranean Coast and as north as Finland. They are often found on bridges, especially near light and over water. The species tends to live on steel objects and is seldom seen on vegetation. Females reach a body length of 10–14mm, and males 8–9mm. Their orb webs can have diameters of up to 70 cm.
The Neriidae are a family of true flies (Diptera) closely related to the Micropezidae. Some species are known as cactus flies, while others have been called banana stalk flies and the family was earlier treated as subfamily of the Micropezidae which are often called stilt-legged flies. Neriids differ from micropezids in having no significant reduction of the fore legs. Neriids breed in rotting vegetation, such as decaying tree bark or rotting fruit. About 100 species are placed in 19 genera. Neriidae are found mainly in tropical regions, but two North American genera occur, each with one species, and one species of Telostylinus occurs in temperate regions of eastern Australia.
Rhagoletis juglandis, also known as the walnut husk fly, is a species of tephritid or fruit fly in the family Tephritidae. It is closely related to the walnut husk maggot Rhagoletis suavis. This species of fly belongs to the R. suavis group, which has a natural history consistent with allopatric speciation. The flies belonging to this group are morphologically distinguishable.
The six-spotted fishing spider is an arachnid from the nursery web spider family Pisauridae. This species is from the genus Dolomedes, or the fishing spiders. Found in wetland habitats throughout North America, these spiders are usually seen scampering along the surface of ponds and other bodies of water. They are also referred to as dock spiders because they can sometimes be witnessed quickly vanishing through the cracks of boat docks. D. triton gets its scientific name from the Greek mythological god Triton, who is the messenger of the big sea and the son of Poseidon.
Empis borealis is a species of dance flies in the fly family Empididae. It is found in most of Europe, except the Balkan Peninsula, the Baltic Region and the Iberian Peninsula. The brown wings of the female are very broad, while the male fly’s wings are narrower.
Empis tessellata is a species of dance fly, in the fly family Empididae. It is included in the subgenus Euempis.
Empis ciliata, the black dance fly, is a species of dance fly, in the fly family Empididae. It is included in the subgenus Euempis.
Diptera is an order of winged insects commonly known as flies. Diptera, which are one of the most successful groups of organisms on Earth, are very diverse biologically. None are truly marine but they occupy virtually every terrestrial niche. Many have co-evolved in association with plants and animals. The Diptera are a very significant group in the decomposition and degeneration of plant and animal matter, are instrumental in the breakdown and release of nutrients back into the soil, and whose larvae supplement the diet of higher agrarian organisms. They are also an important component in food chains.
A nuptial gift is a nutritional gift given by one partner in some animals' sexual reproduction practices.
Pisaurina mira, also known as the American nursery web spiders, is a species of spider in the family Pisauridae. They are often mistaken for wolf spiders (Lycosidae) due to their physical resemblance. P. mira is distinguished by its unique eye arrangement of two rows.
Dryomyza anilis is a common fly from the family Dryomyzidae. The fly is found through various areas in the Northern hemisphere and has brown and orange coloration with distinctive large red eyes. The life span of the fly is not known, but laboratory-reared males can live 28–178 days. D. anilis has recently been placed back in the genus Dryomyza, of which it is the type species. Dryomyzidae were previously part of Sciomyzidae but are now considered a separate family with two subfamilies.
Hylobittacus apicalis is a species of hangingfly in the order Mecoptera, and the only species within the genus Hylobittacus.
Rhamphomyia longicauda, the long-tailed dance fly, is a species of fly commonly found in eastern North America that belongs to the family Empididae and part of the superfamily of dance flies Empidoidea. It is included in the subgenus Rhamphomyia. This species of fly is most known for sex role reversal during courtship, as females put on exaggerated displays and congregate in leks to attract males. Females cannot hunt for food, so they receive protein from nuptial gifts brought to them by males. Female dependence on males for nutrition is the principal cause for sex role reversal in this species of fly.
Argyrodes elevatus, commonly referred to as dew-drop spider, is part of the family Theridiidae that consists of more than 3,000 species. These spiders are most commonly found in subtropical and tropical regions in South and Central America, as well as southern regions of the United States. One of the key distinguishing characteristics of A. elevatus is its kleptoparasitic behavior through which it primarily procures food for survival. Typically 1 or 2 A. elevatus spiders preside in outer areas of webs built by other species of spiders, although it is possible for up to 45 spiders. There are two main mechanisms by which A. elevatus raid the hub of the host’s web to steal insects preyed and wrapped by the host spider. A. elevatus follows an intricate course to the hub of the web to search for prey, using vibrational detection enhanced by laid out threads along the web to find and capture the insect. These spiders are highly efficient, with the theft lasting a maximum of 12 seconds and high success rates. This reliance on a host spider for food has led to adaptations in sleep schedules and alternate food sources to revolve around the host species activity. A. elevatus display a unique courtship routine in which male A. elevatus presents prey wrapped in silk as a nuptial gift to the female spider. The male spider approaches the female, carrying the nuptial gift on its chelicerae while communicating with a distinct courting vibration, followed by copulation. Approximately twenty-four hours after the A. elevatus courtship and copulation series of events, the female spider will lay one to two eggs on the outer regions of the host's web.
The social network of colonies of social insects have been extensively studied as model biological networks.