A mammal is a vertebrate animal of the class Mammalia. Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles and birds, from which their ancestors diverged in the Carboniferous Period over 300 million years ago. Around 6,400 extant species of mammals have been described and divided into 27 orders. The study of mammals is called mammalogy.
Marsupials are a diverse group of mammals belonging to the infraclass Marsupialia. They are natively found in Australasia, Wallacea, and the Americas. One of the defining features of marsupials is their unique reproductive strategy, where the young are born in a relatively undeveloped state and then nurtured within a pouch on their mother's abdomen.
Multituberculata is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, and reached a peak diversity during the Late Cretaceous and Paleocene. They eventually declined from the mid-Paleocene onwards, disappearing from the known fossil record in the late Eocene. They are the most diverse order of Mesozoic mammals with more than 200 species known, ranging from mouse-sized to beaver-sized. These species occupied a diversity of ecological niches, ranging from burrow-dwelling to squirrel-like arborealism to jerboa-like hoppers. Multituberculates are usually placed as crown mammals outside either of the two main groups of living mammals—Theria, including placentals and marsupials, and Monotremata—but usually as closer to Theria than to monotremes. They are considered to be closely related to Euharamiyida and Gondwanatheria as part of Allotheria.
Placental mammals are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placental mammals contains the vast majority of extant mammals, which are partly distinguished from monotremes and marsupials in that the fetus is carried in the uterus of its mother to a relatively late stage of development. The name is something of a misnomer, considering that marsupials also nourish their fetuses via a placenta, though for a relatively briefer period, giving birth to less-developed young, which are then nurtured for a period inside the mother's pouch. Placental mammals represents the only living group within Eutheria, which contains all mammals that are more closely related to placental mammals than they are to marsupials.
Eomaia is a genus of extinct fossil mammals containing the single species Eomaia scansoria, discovered in rocks that were found in the Yixian Formation, Liaoning Province, China, and dated to the Barremian Age of the Lower Cretaceous about 125 million years ago. The single fossil specimen of this species is 10 centimetres (3.9 in) in length and virtually complete. An estimate of the body weight is 20–25 grams (0.71–0.88 oz). It is exceptionally well-preserved for a 125-million-year-old specimen. Although the fossil's skull is squashed flat, its teeth, tiny foot bones, cartilages and even its fur are visible.
Eutheria, also called Pan-Placentalia, is the clade consisting of placentals and all therian mammals that are more closely related to placentals than to marsupials.
Zalambdalestes is an extinct genus of eutherian mammal known from the Upper Cretaceous in Mongolia.
The pouch is a distinguishing feature of female marsupials, monotremes ; the name marsupial is derived from the Latin marsupium, meaning "pouch". This is due to the occurrence of epipubic bones, a pair of bones projecting forward from the pelvis. Marsupials give birth to a live but relatively undeveloped foetus called a joey. When the joey is born it crawls from inside the mother to the pouch. The pouch is a fold of skin with a single opening that covers the teats. Inside the pouch, the blind offspring attaches itself to one of the mother's teats and remains attached for as long as it takes to grow and develop to a juvenile stage.
Akidolestes is an extinct genus of mammals of the family Spalacotheriidae, a group of mammals related to therians.
Mammaliaformes is a clade of synapsid tetrapods that includes the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts during the Late Triassic. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes also includes Docodonta and Hadrocodium.
Tritylodontidae is an extinct family of small to medium-sized, highly specialized mammal-like cynodonts, with several mammalian traits including erect limbs, endothermy, and some details of the skeleton. They were the last-known family of the non-mammaliaform synapsids, persisting into the Early Cretaceous.
The evolution of mammals has passed through many stages since the first appearance of their synapsid ancestors in the Pennsylvanian sub-period of the late Carboniferous period. By the mid-Triassic, there were many synapsid species that looked like mammals. The lineage leading to today's mammals split up in the Jurassic; synapsids from this period include Dryolestes, more closely related to extant placentals and marsupials than to monotremes, as well as Ambondro, more closely related to monotremes. Later on, the eutherian and metatherian lineages separated; the metatherians are the animals more closely related to the marsupials, while the eutherians are those more closely related to the placentals. Since Juramaia, the earliest known eutherian, lived 160 million years ago in the Jurassic, this divergence must have occurred in the same period.
The brushtail possums are the members of the genus Trichosurus in the Phalangeridae, a family of marsupials. They are native to Australia and some small nearby islands. Unique among marsupials, they have shifted the hypaxial muscles from the epipubic to the pelvis, much like in placental mammals, meaning that their breathing cycle is more similar to the latter than to that of other non-eutherian mammals. In general, they are more terrestrially oriented than other possums, and in some ways might parallel primates.
The Phalangeridae are a family of mostly nocturnal marsupials native to Australia, New Guinea, and Eastern Indonesia, including the cuscuses, brushtail possums, and their close relatives. Considered a type of possum, most species are arboreal, and they inhabit a wide range of forest habitats from alpine woodland to eucalypt forest and tropical jungle. Many species have been introduced to various non-native habitats by humans for thousands of years.
Monotremes are mammals of the order Monotremata. They are the only group of living mammals that lay eggs, rather than bearing live young. The extant monotreme species are the platypus and the four species of echidnas. Monotremes are typified by structural differences in their brains, jaws, digestive tract, reproductive tract, and other body parts, compared to the more common mammalian types. Although they are different from almost all mammals in that they lay eggs, like all mammals, the female monotremes nurse their young with milk.
Several mammals are known from the Mesozoic of Madagascar. The Bathonian Ambondro, known from a piece of jaw with three teeth, is the earliest known mammal with molars showing the modern, tribosphenic pattern that is characteristic of marsupial and placental mammals. Interpretations of its affinities have differed; one proposal places it in a group known as Australosphenida with other Mesozoic tribosphenic mammals from the southern continents (Gondwana) as well as the monotremes, while others favor closer affinities with northern (Laurasian) tribosphenic mammals or specifically with placentals. At least five species are known from the Maastrichtian, including a yet undescribed species known from a nearly complete skeleton that may represent a completely new group of mammals. The gondwanathere Lavanify, known from two teeth, is most closely related to other gondwanatheres found in India and Argentina. Two other teeth may represent another gondwanathere or a different kind of mammal. One molar fragment is one of the few known remains of a multituberculate mammal from Gondwana and another has been interpreted as either a marsupial or a placental.
Megaconus is an extinct genus of allotherian mammal from the Middle Jurassic Tiaojishan Formation of Inner Mongolia, China. The type and only species, Megaconus mammaliaformis was first described in the journal Nature in 2013. Megaconus is thought to have been a herbivore that lived on the ground, having a similar posture to modern-day armadillos and rock hyraxes. Megaconus was in its initial description found to be member of a group called Haramiyida. A phylogenetic analysis published along its description suggested that haramiyidans originated before the appearance of true mammals, but in contrast, the later description of the haramiyidan Arboroharamiya in the same issue of Nature indicated that haramyidans were true mammals. If haramiyidans are not mammals, Megaconus would be one of the most basal ("primitive") mammaliaforms to possess fur, and an indicator that fur evolved in the ancestors of mammals and not the mammals themselves. However, later studies cast doubt on the euharamiyidan intrepretation, instead finding it to be a basal allotherian mammal.
Ukhaatherium is a now extinct species of mammal that lived during the upper Cretaceous about 84 to 72 million years ago in today's East Asia. It is known above all from the fossil locality Ukhaa Tolgod, Mongolia. An adult Ukhaatherium has an estimated weight of about 32g and bears several similarities to lipotyphlan insectivorans such as the tenrec.
Marsupionta is a hypothetical clade of mammals containing marsupials and monotremes, but not the placentals. This hypothesis is contrary to the conventional view that marsupials and placentals form a clade (Theria) that excludes monotremes. Marsupionta was proposed in 1947 by the American zoologist William King Gregory and has since been the subject of multiple studies.
Zalambdalestidae is a clade of Asian eutherians occurring during the Cretaceous. Once classified as Glires, features like epipubic bones and various cranial elements have identified these animals as outside of Placentalia, representing thus a specialised clade of non-placental eutherians without any living descendants, and potentially rather different from modern placentals in at least reproductive anatomy.
