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The Formica rufa group is a subgeneric group within the genus Formica , first proposed by William Morton Wheeler. This group contains the mound-building species of Formica commonly termed "wood ants" or "thatch-mound ants", which build prominent nests consisting of a mound of grass, litter, or conifer needles. [1] The species Formica rufa or the red wood ant is the type species of this subgroup.
This particular group can inhabit open deciduous woodlands, dense pine forests, and even moorland. Workers vary in size, the largest reaching about 10 millimetres (0.39 in) long. They can produce formic acid in their abdomens and eject it 12 centimetres (4.7 in) in the air when threatened. [2] The only function of males is to mate in flight with queens. [2] Species previously attributed to the Microgyna group were transferred to the F. rufa group by the Wheelers in 1986.[ citation needed ]
The genetic relationships in F. rufa group colonies can be complex. Colonies can be polygynous (having multiple queens) and these are often polydomous (having multiple nests per colony). Queens may be singly or multiply mated, and in polygynous colonies, may or may not be related. Colonies are rarely, if ever, independently formed. They either bud off from existing colonies or are formed by temporary social parasitism of colonies of F. fusca or other closely related species. A wood ant queen ousts the existing queen, lays eggs, and the F. fusca workers raise her offspring until the nest is taken over. Some species in the F. rufa group sometimes form enormous "supercolonies" consisting of hundreds or thousands of nests. The largest documented example is an F. yessensis colony in Japan covering 2.7 square kilometres (1.0 sq mi) containing an estimated 306 million workers and 1 million queens. [3]
European species include:
North American species include:
Asian species
Formica exsecta is a species of ant found from Western Europe to Asia.
Formica rufibarbis is a European formicine ant of the Formica fusca group. In the classification by Auguste Forel, it is treated in the subgenus Serviformica. F. rufibarbis is subject to a Species Action Plan (SAP) in England, where it is known from only two locations, although it is not considered to be at risk on continental Europe.
Formica is a genus of ants of the family Formicidae, including species commonly known as wood ants, mound ants, thatching ants, and field ants. Formica is the type genus of the Formicidae, and of the subfamily Formicinae. The type species of genus Formica is the European red wood ant Formica rufa. Ants of this genus tend to be between 4 and 8 mm long. Ants belonging to the Formica genus possess a single knob or bump located between their thorax and abdomen. These ants primarily feed on honeydew, a sugary liquid produced by aphids. Formica ants appear to take on a shepherding role with smaller aphids, relocating them to different parts of plants to ensure a continuous food source for the aphids. By doing so, the ants can establish a relatively sustainable honeydew supply for both themselves and their colony.
Formica rufa, also known as the red wood ant, southern wood ant, or horse ant, is a boreal member of the Formica rufa group of ants, and is the type species for that group, being described already by Linnaeus. It is native to Eurasia, with a recorded distribution stretching from the middle of Scandinavia to the northern Iberia and Anatolia, and from Great Britain to Lake Baikal, with unconfirmed reportings of it also to the Russian Far East. There are claims that it can be found in North America, but this is not confirmed in specialised literature, and no recent publication where North American wood ants are listed mentions it as present, while records from North America are all listed as dubious or unconfirmed in a record compilation. Workers head and thorax are colored red and the abdomen brownish-black, usually with a dorsal dark patches on the head and promensonotum, although some individuals may be more uniform reddish and even have some red on the part of the gastern facing the body. In order to separate them from closely related species, specimens needs to be inspected under magnification, where difference in hairyness are among the telling characteristics, with Formica rufa being more hairy than per example Formica polyctena but less hairy than Formica lugubris. Workers are polymorphic, measuring 4.5–9 mm in length. They have large mandibles, and like many other ant species, they are able to spray formic acid from their abdomens as a defence. Formic acid was first extracted in 1671 by the English naturalist John Ray by distilling a large number of crushed ants of this species. These ants primarily eat honeydew from aphids. Some groups form large networks of connected nests with multiple queen colonies, while others have single-queen colonies.
An ant colony is a population of a single ant species able to maintain its complete lifecycle. Ant colonies are eusocial, communal, and efficiently organized and are very much like those found in other social Hymenoptera, though the various groups of these developed sociality independently through convergent evolution. The typical colony consists of one or more egg-laying queens, numerous sterile females and, seasonally, many winged sexual males and females. In order to establish new colonies, ants undertake flights that occur at species-characteristic times of the day. Swarms of the winged sexuals depart the nest in search of other nests. The males die shortly thereafter, along with most of the females. A small percentage of the females survive to initiate new nests.
Carpenter ants are large ants indigenous to many forested parts of the world.
Dorylus, also known as driver ants, safari ants, or siafu, is a large genus of army ants found primarily in central and east Africa, although the range also extends to southern Africa and tropical Asia. The term siafu is a loanword from Swahili, and is one of numerous similar words from regional Bantu languages used by indigenous peoples to describe various species of these ants. Unlike the New World members of the former subfamily Ecitoninae, members of this genus form temporary subterranean bivouacs in underground cavities which they excavate and inhabit - either for a few days or up to three months. Also unlike some New World army ants, driver ants are not specialized predators of other species of ant, instead being more generalistic with a diet consisting of a diversity of arthropods. Colonies are enormous compared to other army ants and can contain over 20 million individuals. As with their American counterparts, workers exhibit caste polymorphism with the soldiers having particularly large heads that power their scissor-like mandibles. They are capable of stinging, but very rarely do so, relying instead on their powerful shearing jaws. Driver ant queens are the largest living ants known, with the largest measuring between 40 - 63 millimeters in total body length depending on their physiological condition.
Formica fusca is a black-colored ant commonly found throughout Europe as well as parts of southern Asia and Africa. It has the common names silky ant or dusky ant. The range within the palaearctic region extends from Portugal in the west to Japan in the east and from Italy in the south to Fennoscandia in the north. Populations from North America have been split off as a separate species, Formica subaenescens. F. fusca nests are usually found in rotten tree stumps or under stones in clearcut areas and along woodland borders and hedgerows.
Crematogaster is an ecologically diverse genus of ants found worldwide, which are characterised by a distinctive heart-shaped gaster (abdomen), which gives them one of their common names, the Saint Valentine ant. Members of this genus are also known as cocktail ants because of their habit of raising their abdomens when alarmed. Most species are arboreal (tree-dwelling). These ants are sometimes known as acrobat ants.
Formica lugubris, also known as the hairy wood ant is commonly found in wooded upland areas across northern Eurasia. Colonies construct large thatched mound nests occupied by thousands of workers, and one or more queens. Workers look similar to other species of wood ants, but Formica lugubris workers can be identified by a fringe of hairs that reaches down to their eyes and prominent hairs between the facets of their compound eyes. Workers can reach sizes of up to 9 mm long; queens are larger, reaching 12 mm long.
Formica polyctena is a species of European red wood ant in the genus Formica and large family Formicidae. The species was first described by Arnold Förster in 1850. The latin species name polyctena is from Greek and literally means 'many cattle', referring to the species' habit of farming aphids for honeydew food. It is found in many European countries. It is a eusocial species, that has a distinct caste system of sterile workers and a very small reproductive caste. The ants have a genetic based cue that allow them to identify which other ants are members of their nest and which are foreign individuals. When facing these types of foreign invaders the F. polyctena has a system to activate an alarm. It can release pheromones which can trigger an alarm response in other nearby ants.
Formica pratensis, the black-backed meadow ant, is a species of European red wood ant in the family Formicidae.
Formica sanguinea, or blood-red ant, is a species of facultative slave-maker ant in the genus Formica characterized by the ability to secrete formic acid. It ranges from Central and Northern Europe through Russia to Japan, China, the Korean Peninsula, Africa and also the United States. This species is coloured red and black with workers up to 7 mm long.
Worker policing is a behavior seen in colonies of social hymenopterans whereby worker females eat or remove eggs that have been laid by other workers rather than those laid by a queen. Worker policing ensures that the offspring of the queen will predominate in the group. In certain species of bees, ants and wasps, workers or the queen may also act aggressively towards fertile workers. Worker policing has been suggested as a form of coercion to promote the evolution of altruistic behavior in eusocial insect societies.
Formica cinerea is a species of ant in the family Formicidae.
Formica truncorum is a species of wood ant from the genus Formica. It is distributed across a variety of locations worldwide, including central Europe and Japan. Workers can range from 3.5 to 9.0mm and are uniquely characterized by small hairs covering their entire bodies. Like all other ants, F. truncorum is eusocial and demonstrates many cooperative behaviors that are unique to its order. Colonies are either monogynous, with one queen, or polygynous, with many queens, and these two types of colonies differ in many characteristics.
Formica incerta is a species of ant found in eastern North America. It is the most common species of Formica in many areas, and excavates underground nests with small entrance holes. Its diet includes nectar produced by extrafloral nectaries and honeydew, which it obtains from aphids and treehoppers. It is the main host for the slave-making ant Polyergus lucidus. F. incerta was first described by Italian entomologist Carlo Emery in 1893. Its specific name comes from the Latin incertus meaning "uncertain" and seems particularly apt given the subsequent uncertainty as to the validity of the species and the difficulty in distinguishing this ant from other species living in the same area.
An ant supercolony is an exceptionally large ant colony, consisting of a high number of spatially separated but socially connected nests of a single ant species, spread over a large area without territorial borders. Supercolonies are typically polygynous, containing many egg-laying females. Workers and queens from different nests within the same supercolony can freely move among the nests, and all workers cooperate indiscriminately with each other in collecting food and care of the brood, and show no apparent mutual aggressive behavior.