The Carboniferous is a geologic period and system of the Paleozoic that spans 60 million years from the end of the Devonian Period 358.9 million years ago (mya), to the beginning of the Permian Period, 298.9 mya. The name Carboniferous means "coal-bearing", from the Latin carbō ("coal") and ferō, and refers to the many coal beds formed globally during that time.
Ammonoids are a group of extinct marine mollusc animals in the subclass Ammonoidea of the class Cephalopoda. These molluscs, commonly referred to as ammonites, are more closely related to living coleoids than they are to shelled nautiloids such as the living Nautilus species. The earliest ammonites appeared during the Devonian, with the last species vanishing during or soon after the Cretaceous–Paleogene extinction event.
The Nautilida constitute a large and diverse order of generally coiled nautiloid cephalopods that began in the mid Paleozoic and continues to the present with a single family, the Nautilidae which includes two genera, Nautilus and Allonautilus, with six species. All told, between 22 and 34 families and 165 to 184 genera have been recognised, making this the largest order of the subclass Nautiloidea.
Goniatids, informally goniatites, are ammonoid cephalopods that form the order Goniatitida, derived from the more primitive Agoniatitida during the Middle Devonian some 390 million years ago. Goniatites (goniatitids) survived the Late Devonian extinction to flourish during the Carboniferous and Permian only to become extinct at the end of the Permian some 139 million years later.
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day.
The Bactritida are a small order of more or less straight-shelled (orthoconic) cephalopods that first appeared during the Emsian stage of the Devonian period with questionable origins in Pragian stage before 409 million years ago, and persisted until Carnian pluvial event in the upper middle Carnian stage of the Triassic period. They are considered ancestors of the ammonoids, as well as of the coleoids.
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician to the Middle Devonian. Some, such as Aphetoceras and Estonioceras, are loosely coiled and gyroconic; others, such as Campbelloceras, Tarphyceras, and Trocholites, are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular, as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably Bassleroceras, around the end of the Gasconadian through forms like Aphetoceras. Close coiling developed rather quickly, and both gyroconic and evolute forms are found in the early middle Canadian.
Beekmanoceras is a small cephalopod from the Middle Canadian Epoch of New York with a loosely coiled, gyroconic, shell in which the whorls are not in contact and the siphuncle is on the inner or concave side of the whorl. Furnish and Glenister (1964) placed Beekmanoceras in the Trocholitidae (Tarphycerida), interpreting the curvature to be ventral side convex, i.e. exogastastric and the siphuncle to be dorsal. Flower (1964) included Beekmanoceras in the Ellesmeroceratidae believing the siphuncle to be ventral and the curvature to be endogastric with the ventral side concave.
Osbornoceras is a genus of Lower Silurian cyrtoconic nautiloid cephalopods known from Ohio and possibly Manitoba, one of five general currently included in the oncocerid family Karoceratidae.
The Tainoceratoidea are a superfamily in the cephalopod order Nautilida characterized by straight to loosely coiled shells, generally to a degree such that the width is greater than the height, to quadrate whorl section. Many bore spines, ribs, frills, wings, or nodes. In early forms, the siphuncle is generally near ventral, but more variable in advanced forms.
The Columbus Limestone is a mapped bedrock unit consisting primarily of fossiliferous limestone, and it occurs in Ohio, Pennsylvania, and Virginia in the United States, and in Ontario, Canada.
The Naedyceras group comprises three similar and closely related openly coiled, gyroconic, genera within oncocerid family, Brevicoceratidae: Naedyceras, Gonionaedyceras, and Gyronaedyceras.
Rhynchonelliformea is a major subphylum and clade of brachiopods. It is roughly equivalent to the former class Articulata, which was used previously in brachiopod taxonomy up until the 1990s. These so-called articulated brachiopods have many anatomical differences relative to "inarticulate" brachiopods of the subphyla Linguliformea and Craniformea. Articulates have hard calcium carbonate shells with tongue-and-groove hinge articulations and separate sets of simple opening and closing muscles.
Sceptrites is an orthoceratoid genus contained within the pseudorthocerid family Spyroceratidae known by its slightly compressed, possibly endogastrically curved, cyrtoconic shell. Sceptrites has been found in Devonian sediments in Ohio and N.Y. in the United States and in Victoria, Australia.
Trochoceras is a genus of trochoidally coiled nautiloid cephalopods placed in the nautilid family Rutoceratidae that lived during the Middle and Late Devonian in what is now central Europe.
The Nephriticeratidae is a family of early Paleozoic nautilod cephalopods included in the Barrandeocerina, distinguished by mostly cyrtoconic as well as gyroconic, sepenticontic, and sinstrally torticonic shells with large, typically straight necked (orthchoanitic) siphuncles. As for the suborder, connecting rings are thin.
Gonatocyrtoceras is a genus of oncocerid nautiloids belonging to the family Acleistoceratidae, that lived during the Middle Devonian. Gonatocyrtoceras is characterized by strongly curved, depressed, breviconic shells. Curvature is exogastric, meaning the lower, or ventral, side is convex in longitudinal profile. The aperture is without a hyponomic sinus. The siphuncle is small and subventral.
Discosoridae comprise a family of endogastric discosorids, with endocones in the siphuncle, ranging from the Middle Silurian to Middle Devonian.
Diademoceras is a genus of nautiloid cephalopods from the middle Devonian of North America, named by Rousseau Flower in 1945. The genus is a tainoceratacean included in the nautilid family Rutoceratidae.
