Holmiceras Temporal range: late L Ordovician (Arenig) | |
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Scientific classification | |
Kingdom: | Animalia |
Phylum: | Mollusca |
Class: | Cephalopoda |
Subclass: | Nautiloidea |
Order: | † Tarphycerida |
Family: | † Lituitidae |
Genus: | † Holmiceras Hyatt, 1894 |
Holmiceras is one of two ancestral lituitids from the late Early Ordovician (Arenigian). The other being Ancistroceras .
Holmiceras begins with about 1.5 to 2 loosely coiled whorls followed by a rapidly expanding orthocone, much like Ancistroceras except for the apical whorls and that Holmiceras is known only from the late Early Ordovician whereas Ancestroceras extends through the middle of the period.
Holmiceras may have given rise to Angelinoceras which begins with an expanding open spiral.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
Trocholites is a tarphycerid genus in the family Trocholitidae from the Middle and Late Ordovician with a gradually expanding, weakly ribbed shell; whorls in contact, dorsum slightly impressed; cross section depressed, venter and sides rounded; siphuncle close to but not at the dorsal margin.
The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian, in which the connecting rings are thin and siphuncle segments are variably expanded. At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three.
The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician to the Middle Devonian. Some, such as Aphetoceras and Estonioceras, are loosely coiled and gyroconic; others, such as Campbelloceras, Tarphyceras, and Trocholites, are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular, as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably Bassleroceras, around the end of the Gasconadian through forms like Aphetoceras. Close coiling developed rather quickly, and both gyroconic and evolute forms are found in the early middle Canadian.
Curtoceras is a genus in the tarphycerid family Trocholitidae found widespread in the late Early and Middle Ordovician of North America and northern Europe. Curtoceras has a shell that is gradually expanded, with half the fully mature body chamber divergent from the preceding volution. Whorl sections are near equidimentional with the inner margin (dosum) moderately impressed. The surface may be smooth or weakly ribbed. The siphuncle is ventral in the initial chamber and becomes dorsal after one volution. With the exception of the dorsal siphuncle, Curtoceras is somewhat similar to the tarphyceratid Campbelloceras
The Tarphyceratidae are tightly coiled, evolute Tarphycerida with ventral siphuncles. The dorsum is characteristically impressed where the whorl presses against the venter of the previous. The Tarphyceratidae are derived from Bassleroceras or possibly from some member of the Estonioceratidae.
The Trocholitidae are Tarphycerida with whorls in close contact as with the Tarphyceratidae, but in which the siphuncle, similar in structure, becomes dorsal. The Trocholitidae are derived from the Tarphyceratidae, perhaps from different tarphyceratids.
Pseudorthocerida is an order of generally straight longiconic Orthoceratoids with a subcentral to marginal cyrtochoanitic siphuncle composed of variably expanded segments which may contain internal deposits that may develop into a continuous parietal lining.. Cameral deposits are common and concentrated ventrally. Apices typically have a slight to moderate exogastric curvature
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.
Orthoceratoidea is a subclass, formerly considered an infraclass or a superorder, that comprises Cephalopoda orders that have orthoconic to slightly cyrtoconic shells and central to subcentral siphuncles in which there may be internal deposits. Currently, Orthoceratoidea comprises the orders Dissidocerida, Ascocerida, Pseudorthocerida, Lituitida and Orthocerida.
Oncoceratidae is a family of nauatiloid cephalopods in the order Oncocerida established by Hyatt, 1884, that range from the Middle Ordovician to the Upper Silurian.
The Bassleroceratidae is a family of gradually expanding, smooth ellesmerocerids with a slight to moderate exogastric curvature, subcircular to strongly compressed cross section, and ventral orthochaonitc siphuncle. The ventral side is typically more sharply rounded than the dorsal side and septa are close spaced. Connecting rings are thick and slightly expanded into the siphuncle, making the segments slightly concave; characteristic of the Ellesmerocerida.
The Tripteroceratidae is a family of depressed, straight to slightly curved nautiloid cephalopods from the middle and upper Ordovician with generally flattened venters and empty siphuncles with straight to inflated segments included in the Oncocerida.
Ancistroceras is one of the two ancestral lituitids from the late Early Ordovician (Arenigian). The other being Holmiceras.
The Lituitidae are a family of evolved tarphycerids characterized by a long orthoconic section that follows a coiled juvenile portion at the apex, along with a generally tubular siphuncle, which like that of the barrandeocerids is composed of thin connecting rings.
Angelinoceras is a genus of lituitids from the Middle Ordovician that starts off with an open spiral of about 1.5 strongly compressed whorls followed by a straight orthoconic section that continues to expand for a length about equal to the diameter of the coiled portion before retaining a more or less consistent diameter. The juvenile spiral portion has a deep indentation on the ventral side of the aperture for the water-jet funnel, known as a hyponomic sinus, that becomes broad and shallow in the straight-shelled adult portion.
Rhynchorthoceras is a Middle Ordovician genus characterized by a rapidly expanded, weakly annulate orthocone, like the orthoconic section of Ancistroceras, but with only a curved, cyrtoconic apex instead of juvenile whorls.
Barrandeoceras is a large, coiled, Middle Ordovician nautiloid cephalopod and part of the Tarphycerida. The shell is serpenticonic with whorls touching but not embracing. The adult body chamber becomes freed of the preceding whorl, a rather common character among tarphyceroids. Whorl section is oval, somewhat more narrowly rounded ventrally, on the outer rim, than dorsally, on the inner rim. Prominent lateral ribs, at least on inner whorls. Grow lines show a distinct hyponomic sinus. Sutures have lateral lobes. The siphuncle is subcentral.
Plectoceras is a genus of nautiloids included in the tarphycerid suborder Barrandeocerina that lived during the Middle and Late Ordovician. It has been found widespread in the Middle and Upper Ordovician of North America.
Aethoceras is a genus of Tarphycerida nautiloids included in the family Estonioceratidae for which the shell is a loosely coiled, gradually expanding dextral torticone with a slightly depressed whorl section. Siphuncle small, ventral, submarginal. Whorl section somewhat resembles early stages of Estonioceras in being laterally fanged. Trochoidal coiling brings to mind the later Trocholitidae. Aethoceras was originally found in the lower Ordovician of Western Australia.