Human evolution is the evolutionary process within the history of primates that led to the emergence of Homo sapiens as a distinct species of the hominid family that includes all the great apes. This process involved the gradual development of traits such as human bipedalism, dexterity, and complex language, as well as interbreeding with other hominins, indicating that human evolution was not linear but weblike. The study of the origins of humans involves several scientific disciplines, including physical and evolutionary anthropology, paleontology, and genetics; the field is also known by the terms anthropogeny, anthropogenesis, and anthropogony.
Early modern human (EMH), or anatomically modern human (AMH), are terms used to distinguish Homo sapiens that are anatomically consistent with the range of phenotypes seen in contemporary humans, from extinct archaic human species. This distinction is useful especially for times and regions where anatomically modern and archaic humans co-existed, for example, in Paleolithic Europe. Among the oldest known remains of Homo sapiens are those found at the Omo-Kibish I archaeological site in south-western Ethiopia, dating to about 233,000 to 196,000 years ago, the Florisbad site in South Africa, dating to about 259,000 years ago, and the Jebel Irhoud site in Morocco, dated about 315,000 years ago.
Homo is a genus of great ape that emerged from the genus Australopithecus and encompasses only a single extant species, Homo sapiens, along with a number of extinct species classified as either ancestral or closely related to modern humans; these include Homo erectus and Homo neanderthalensis. The oldest member of the genus is Homo habilis, with records of just over 2 million years ago. Homo, together with the genus Paranthropus, is probably most closely related to the species Australopithecus africanus within Australopithecus. The closest living relatives of Homo are of the genus Pan, with the ancestors of Pan and Homo estimated to have diverged around 5.7-11 million years ago during the Late Miocene.
Paleoanthropology or paleo-anthropology is a branch of paleontology and anthropology which seeks to understand the early development of anatomically modern humans, a process known as hominization, through the reconstruction of evolutionary kinship lines within the family Hominidae, working from biological evidence and cultural evidence.
The Mousterian is an archaeological industry of stone tools, associated primarily with the Neanderthals in Europe, and to the earliest anatomically modern humans in North Africa and West Asia. The Mousterian largely defines the latter part of the Middle Paleolithic, the middle of the West Eurasian Old Stone Age. It lasted roughly from 160,000 to 40,000 BP. If its predecessor, known as Levallois or Levallois-Mousterian, is included, the range is extended to as early as c. 300,000–200,000 BP. The main following period is the Aurignacian of Homo sapiens.
The Aterian is a Middle Stone Age stone tool industry centered in North Africa, from Mauritania to Egypt, but also possibly found in Oman and the Thar Desert. The earliest Aterian dates to c. 150,000 years ago, at the site of Ifri n'Ammar in Morocco. However, most of the early dates cluster around the beginning of the Last Interglacial, around 150,000 to 130,000 years ago, when the environment of North Africa began to ameliorate. The Aterian disappeared around 20,000 years ago.
The Max Planck Institute for Evolutionary Anthropology is a research institute based in Leipzig, Germany, that was founded in 1997. It is part of the Max Planck Society network.
Neanderthals became extinct around 40,000 years ago. Hypotheses on the causes of the extinction include violence, transmission of diseases from modern humans which Neanderthals had no immunity to, competitive replacement, extinction by interbreeding with early modern human populations, natural catastrophes, climate change and inbreeding depression. It is likely that multiple factors caused the demise of an already low population.
Homo rhodesiensis is the species name proposed by Arthur Smith Woodward (1921) to classify Kabwe 1, a Middle Stone Age fossil recovered from Broken Hill mine in Kabwe, Northern Rhodesia. In 2020, the skull was dated to 324,000 to 274,000 years ago. Other similar older specimens also exist.
The Omo remains are a collection of hominin bones discovered between 1967 and 1974 at the Omo Kibish sites near the Omo River, in Omo National Park in south-western Ethiopia. The bones were recovered by a scientific team from the Kenya National Museums directed by Richard Leakey and others. The remains from Kamoya's Hominid Site (KHS) were called Omo I and those from Paul I. Abell's Hominid Site (PHS) were called Omo II.
Human taxonomy is the classification of the human species within zoological taxonomy. The systematic genus, Homo, is designed to include both anatomically modern humans and extinct varieties of archaic humans. Current humans have been designated as subspecies Homo sapiens sapiens, differentiated, according to some, from the direct ancestor, Homo sapiens idaltu.
Archaic humans is a broad category denoting all species of the genus Homo that are not Homo sapiens. Among the earliest modern human remains are those from Jebel Irhoud in Morocco, Florisbad in South Africa (259 ka), Omo-Kibish I in southern Ethiopia, and Apidima Cave in Southern Greece. Some examples of archaic humans include H. antecessor (1200–770 ka), H. bodoensis (1200–300 ka), H. heidelbergensis (600–200 ka), Neanderthals, H. rhodesiensis (300–125 ka) and Denisovans.
Jebel Irhoud or Adrar n Ighoud, is an archaeological site located just north of the town of Tlet Ighoud in Youssoufia Province, approximately 50 km (30 mi) south-east of the city of Safi in Morocco. It is noted for the hominin fossils that have been found there since the discovery of the site in 1960. Originally thought to be Neanderthals, the specimens have since been assigned to Homo sapiens and, as reported in 2017, have been dated to roughly 300,000 years ago.
The Florisbad Skull is an important human fossil of the early Middle Stone Age, representing either late Homo heidelbergensis or early Homo sapiens. It was discovered in 1932 by T. F. Dreyer at the Florisbad site, Free State Province, South Africa.
Scladina, or Sclayn Cave, is an archaeological site located in Wallonia in the town of Sclayn, in the Andenne hills in Belgium, where excavations since 1978 have provided the material for an exhaustive collection of over thirteen thousand Mousterian stone artifacts and the fossilized remains of an especially ancient Neanderthal, called the Scladina child were discovered in 1993.
Katerina Harvati is a Greek paleoanthropologist and expert in human evolution. She specializes in the broad application of 3-D geometric morphometric and virtual anthropology methods to paleoanthropology. Since 2009, she is full professor and director of Paleoanthropology at the University of Tübingen, Germany. From 2020 to 2023 she was Director of the Institute for Archaeological Sciences and since 2023 she is Director of the Senckenberg Centre for Human Evolution and Palaeoenvironment at the University of Tübingen.
The Xiahe mandible is a hominin fossil jaw (mandible) discovered in Baishiya Karst Cave, located on the northeastern edge of the Tibetan Plateau in Xiahe County, Gansu, China. By the use of palaeoproteomic analysis, it is the first confirmed discovery of a Denisovan fossil outside of Denisova Cave, and the most complete confirmed Denisovan fossil. This fossil discovery shows that archaic hominins were present in a high-altitude, low-oxygen environment around 160,000 years ago. Discover, Science News and Nova all named the discovery of the mandible in their lists of Top Science Stories of 2019.
Apidima Cave is a complex of five caves located on the western shore of Mani Peninsula in southern Greece. A systematic investigation of the cave has yielded Neanderthal and Homo sapiens fossils from the Palaeolithic era.
The Salé cranium is a pathological specimen of enigmatic Middle Pleistocene hominin discovered from Salé, Morocco by quarrymen in 1971. Since its discovery, the specimen has variously been classified as Homo sapiens, Homo erectus, Homo rhodesiensis/bodoensis, or Homo heidelbergensis. Its pathological condition and mosaic anatomy has proved difficult to classify. It was discovered with few faunal fossils and no lithics, tentatively dated to 400 ka by some sources.
