List of Ice Age species preserved as permafrost mummies

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Yuka - a woolly mammoth mummy on display in Moscow Mamontionok Iuka.JPG
Yuka – a woolly mammoth mummy on display in Moscow

This is a list of Ice Age species preserved as permafrost mummies. It includes all known species that have had their tissues partially preserved within the permafrost layer of the Arctic and Subarctic. Most went extinct during the Late Pleistocene extinctions while some are still extant today. They have been listed to the most specific known taxonomic rank.

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Permafrost mummies provide crucial insights into the physiology and life histories of Pleistocene organisms, due to how well the preservation process keeps the specimens from decomposing. The constant presence of permafrost is able to preserve the soft tissues of organisms through a process similar to freeze-drying. [1] With such complete preservation of tissues, it is possible to determine numerous things from the such as: DNA, eDNA, [2] evolutionary history, [3] gut contents, [4] and trophic dynamics. [5] Studies have even shown that the process is so complete there is evidence of nucleic activity. [6]

Some of these specimens are on display at the Kingdom of the Permafrost museum near Yakutsk. [7]

(E) - denote an extinct species

Artiodactyla (Even-toed ungulates)

Bovidae (Bovines)

Cervidae (Deer)

Carnivora

Canidae (Dogs)

Felidae (Cats)

Mustelidae

Ursidae (Bears)

Lagomorpha

Ochotonidae (Pika)

Leporidae (Rabbits and hares)

Perissodactyla (Odd-toed ungulates)

Equidae (Horses)

Rhinocerotidae (Rhinoceroses)

Proboscidea

Elephantidae (Elephants)

Rodentia (Rodents)

Cricetidae

Sciuridae (Squirrels)

Passeriformes (Perching Birds)

Alaudidae (Larks)

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<span class="mw-page-title-main">Bison</span> Genus of mammals

A bison is a large bovine in the genus Bison within the tribe Bovini. Two extant and numerous extinct species are recognised.

<span class="mw-page-title-main">Mammoth</span> Extinct genus of mammals

A mammoth is any species of the extinct elephantid genus Mammuthus. They lived from the late Miocene epoch into the Holocene about 4,000 years ago, and various species existed in Africa, Europe, Asia, and North America. Mammoths are distinguished from living elephants by their spirally twisted tusks and in at least some later species, the development of numerous adaptions to living in cold environments, including a thick layer of fur.

<span class="mw-page-title-main">Pleistocene</span> First epoch of the Quaternary Period

The Pleistocene is the geological epoch that lasted from c. 2.58 million to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed in 2009 by the International Union of Geological Sciences, the cutoff of the Pleistocene and the preceding Pliocene was regarded as being 1.806 million years Before Present (BP). Publications from earlier years may use either definition of the period. The end of the Pleistocene corresponds with the end of the last glacial period and also with the end of the Paleolithic age used in archaeology. The name is a combination of Ancient Greek πλεῖστος (pleîstos), meaning "most", and καινός, meaning "new".

<span class="mw-page-title-main">Beringia</span> Geographic region of Asia and North America currently partly submerged

Beringia is defined today as the land and maritime area bounded on the west by the Lena River in Russia; on the east by the Mackenzie River in Canada; on the north by 72 degrees north latitude in the Chukchi Sea; and on the south by the tip of the Kamchatka Peninsula. It includes the Chukchi Sea, the Bering Sea, the Bering Strait, the Chukchi and Kamchatka Peninsulas in Russia as well as Alaska in the United States and the Yukon in Canada.

<span class="mw-page-title-main">Woolly rhinoceros</span> Extinct species of rhinoceros of northern Eurasia

The woolly rhinoceros, simply known as woolly rhino, is an extinct species of rhinoceros that inhabited northern Eurasia during the Pleistocene epoch. The woolly rhinoceros was a member of the Pleistocene megafauna. The woolly rhinoceros was covered with long, thick hair that allowed it to survive in the extremely cold, harsh mammoth steppe. It had a massive hump reaching from its shoulder and fed mainly on herbaceous plants that grew in the steppe. Mummified carcasses preserved in permafrost and many bone remains of woolly rhinoceroses have been found. Images of woolly rhinoceroses are found among cave paintings in Europe and Asia. The species range contracted towards Siberia beginning around 17,000 years ago, with the youngest known records being around 14,000 years old in northeast Siberia, coinciding with the Bølling–Allerød warming, which likely disrupted its habitat.

<span class="mw-page-title-main">Steppe bison</span> Extinct species of mammal

The steppe bison or steppe wisent is an extinct species of bison. It was widely distributed across the mammoth steppe, ranging from Western Europe to eastern Beringia in North America during the Late Pleistocene. It is ancestral to all North American bison, including ultimately modern American bison. Three chronological subspecies, Bison priscus priscus, Bison priscus mediator, and Bison priscus gigas, have been suggested.

<i>Equus</i> (genus) Genus of mammals which includes horses, donkeys, and zebras

Equus is a genus of mammals in the family Equidae, which includes horses, asses, and zebras. Within the Equidae, Equus is the only recognized extant genus, comprising seven living species. Like Equidae more broadly, Equus has numerous extinct species known only from fossils. The genus most likely originated in North America and spread quickly to the Old World. Equines are odd-toed ungulates with slender legs, long heads, relatively long necks, manes, and long tails. All species are herbivorous, and mostly grazers, with simpler digestive systems than ruminants but able to subsist on lower-quality vegetation.

<i>Panthera spelaea</i> Extinct species of lion

Panthera spelaea, also known as the cave lion or steppe lion, is an extinct Panthera species that most likely evolved in Europe after the third Cromerian interglacial stage, less than 600,000 years ago. Genetic analysis of ancient DNA has revealed that while closely related, it was a distinct species genetically isolated from the modern lion occurring in Africa and Asia, with the genetic divergence between the two species variously estimated between 1.9 million and 600,000 years ago. It is closely related and probably ancestral to the American lion. The species ranged from Western Europe to eastern Beringia in North America, and was a prominent member of the mammoth steppe fauna. It became extinct about 13,000 years ago.

<span class="mw-page-title-main">Mammoth steppe</span> Prehistoric biome

During the Last Glacial Maximum, the mammoth steppe, also known as steppe-tundra, was once the Earth's most extensive biome. It stretched east-to-west, from the Iberian Peninsula in the west of Europe, across Eurasia to North America, through Beringia and Canada; from north-to-south, the steppe reached from the arctic islands southward to China. The mammoth steppe was cold and dry, and relatively featureless, though topography and geography varied considerably throughout. Some areas featured rivers which, through erosion, naturally created gorges, gulleys, or small glens. The continual glacial recession and advancement over millennia contributed more to the formation of larger valleys and different geographical features. Overall, however, the steppe is known to be flat and expansive grassland. The vegetation was dominated by palatable, high-productivity grasses, herbs and willow shrubs.

<span class="mw-page-title-main">Spy Cave</span> Archaeological site in Belgium

Spy Cave is located in Wallonia near Spy in the municipality of Jemeppe-sur-Sambre, Namur Province, Belgium above the left bank of the Orneau River. Classified as a premier Heritage site of the Walloon Region, the location ranks among the most significant paleolithic sites in Europe. The cave consists of numerous small chambers and corridors.

<span class="mw-page-title-main">Ancient DNA</span> Method of archaeological study

Ancient DNA (aDNA) is DNA isolated from ancient specimens. Due to degradation processes ancient DNA is more degraded in comparison with contemporary genetic material. Even under the best preservation conditions, there is an upper boundary of 0.4–1.5 million years for a sample to contain sufficient DNA for sequencing technologies. The oldest sample ever sequenced is estimated to be 1.65 million years old. Genetic material has been recovered from paleo/archaeological and historical skeletal material, mummified tissues, archival collections of non-frozen medical specimens, preserved plant remains, ice and from permafrost cores, marine and lake sediments and excavation dirt. In 2022, two-million year old genetic material was found in Greenland, and is currently considered the oldest DNA discovered so far.

<span class="mw-page-title-main">European wild ass</span> Extinct species of mammal

The European wild ass or hydruntine is an extinct equine from the Middle Pleistocene to Late Holocene of Europe and West Asia, and possibly North Africa. It is a member of the subgenus Asinus, and closely related to the living Asiatic wild ass. The specific epithet, hydruntinus, means from Otranto.

<i>Bison occidentalis</i> Extinct species of mammal

Bison occidentalis is an extinct species of bison that lived in North America, from about 11,700 to 5,000 years ago, spanning the end of the Pleistocene to the mid-Holocene.

<span class="mw-page-title-main">Woolly mammoth</span> Extinct species of mammoth from the Quaternary period

The woolly mammoth is an extinct species of mammoth that lived from the Middle Pleistocene until its extinction in the Holocene epoch. It was one of the last in a line of mammoth species, beginning with the African Mammuthus subplanifrons in the early Pliocene. The woolly mammoth began to diverge from the steppe mammoth about 800,000 years ago in Siberia. Its closest extant relative is the Asian elephant. The Columbian mammoth lived alongside the woolly mammoth in North America, and DNA studies show that the two hybridised with each other.

<span class="mw-page-title-main">Late Pleistocene extinctions</span> Extinctions of large mammals in the Late Pleistocene

The Late Pleistocene to the beginning of the Holocene saw numerous extinctions of predominantly megafaunal animal species, which resulted in a collapse in faunal density and diversity across the globe. The extinctions during the Late Pleistocene are differentiated from previous extinctions by the widespread absence of ecological succession to replace these extinct megafaunal species, and the regime shift of previously established faunal relationships and habitats as a consequence. The timing and severity of the extinctions varied by region and are thought to have been driven by varying combinations of human and climatic factors. Human impact on megafauna populations is thought to have been driven by hunting ("overkill"), as well as possibly environmental alteration. The relative importance of human vs climatic factors in the extinctions has been the subject of long-running controversy.

<i>Arctodus</i> Extinct genus of bears

Arctodus is an extinct genus of short-faced bear that inhabited North America during the Pleistocene. There are two recognized species: the lesser short-faced bear and the giant short-faced bear. Of these species, A. simus was larger, is known from more complete remains, and is considered one of the most charismatic of North America's megafauna. A. pristinus was largely restricted to the Early Pleistocene of the Eastern United States, whereas A. simus had a broader range, with most finds being from the Late Pleistocene of the United States, Mexico and Canada. A. simus evolved from A. pristinus, but both species likely overlapped in the Middle Pleistocene. Both species are relatively rare in the fossil record.

<span class="mw-page-title-main">Yuka (mammoth)</span> Mummified mammoth carcass

Yuka is the best-preserved woolly mammoth carcass ever found. It was discovered by local Siberian tusk hunters in August 2010. They turned it over to local scientists, who made an initial assessment of the carcass in 2012. It is displayed in Moscow.

<i>Homo longi</i> Archaic human from China, 146,000 BP

Homo longi is an extinct species of archaic human identified from a nearly complete skull, nicknamed 'Dragon Man', from Harbin on the Northeast China Plain, dating to at minimum 146,000 years ago during the Middle Pleistocene. The skull was discovered in 1933 along the Songhua River while the Dongjiang Bridge was under construction for the Manchukuo National Railway. Due to a tumultuous wartime atmosphere, it was hidden and only brought to paleoanthropologists in 2018. H. longi has been suggested to be the same species as the Denisovans, though this is unconfirmable without genetic testing.

Equus lenensis, the Lena horse, is an extinct species of horse from the Late Pleistocene and Holocene of Siberia, Some sources have considered it a subspecies of the wild horse. Genetic studies show that E. lenensis does not descend from the last common ancestor of living horses, and is estimated to have diverged from them approximately 115,000 years ago, though it is more closely related to modern horses than either are to a highly divergent horse lineage from the Holocene of the Iberian Peninsula. The youngest remains of the species date to 5,000 years Before Present.

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