This is a list of animals that markedly feature external asymmetry in some form. They are exceptions to the general pattern of symmetry in biology. In particular, these animals do not exhibit bilateral symmetry which permits streamlining and is common in animals. [1]
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The crossbill has an unusual beak in which the upper and lower tips cross each other. [2]
The wrybill is the only species of bird with a beak that is bent sideways (always to the right). [2]
Many owl species, such as the barn owl, have asymmetrically positioned ears that enhance sound positioning. [3]
Many flatfish, such as flounders, have eyes placed asymmetrically in the adult fish. The fish has the usual symmetrical body structure when it is young, but as it matures and moves to living close to the sea bed, the fish lies on its side, and the head twists so that both eyes are on the top. [5]
The jaws of the scale-eating cichlid Perissodus microlepis occur in two distinct morphological forms. One morph has its jaw twisted to the left, allowing it to eat scales more readily on its victim’s right flank. The other morph has its jaw twisted to the right, which makes it easier to eat scales on its victim’s left flank. The relative abundance of the two morphs in populations is regulated by frequency-dependent selection. [4] [6] [7]
The narwhal has a helical tusk on its upper left jaw. Odobenocetops , an extinct toothed whale, may have possessed similar asymmetrical dentition, though it differed from the narwhal in possessing two erupted, rear-facing tusks with the right significantly longer than the left. [8]
The sperm whale (Physeter macrocephalus) has a single nostril on its upper left head. The right nostril forms a phonic lip. The source of the air forced through the phonic lips is the right nasal passage. While the left nasal passage opens to the blow hole, the right nasal passage has evolved to supply air to the phonic lips. It is thought that the nostrils of the land-based ancestor of the sperm whale migrated through evolution to their current functions, the left nostril becoming the blowhole and the right nostril becoming the phonic lips.[ citation needed ]
The fin whale (Balaenoptera physalus) has complex and asymmetrical coloration on its head, with the jaw dark grey on one side and white on the other. [9]
The caribou or reindeer (Rangifer tarandus) has asymmetrical antlers. Adult males, in particular, usually possess one brow tine formed into a "shovel" shape. [10] Suggestions for its function include eye protection during antler-threshing courtship displays [11] or as an offensive weapon during the mating season. [12]
Honey badgers of the subspecies signata have a second lower molar on the left side of their jaws, but not the right. [13]
Humans show a systematic aurofacial asymmetry, meaning that the face (eyes, nose and mouth) are displaced to the left with respect to the midplane between the ears. In young children this asymmetry is on average 4 degrees and is easily recognized (See also: Axial Twist theory). [14]
Iwasaki's snail-eater snake (Pareas iwasakii) is a snail-eating specialist; even newly hatched individuals feed on snails. It has asymmetric jaws, which facilitates feeding on snails with dextral (clockwise coiled) shells. A consequence of this asymmetry is that this snake is much less adept at preying on sinistral (counterclockwise coiled) snails. [15] [16] [17] [18] [19] [20] [21] [22]
Fiddler crabs and hermit crabs have one claw much larger than the other, which in hermit crabs is used as a makeshift "door" to block the opening of the shell when it retracts inside. If a male fiddler loses its large claw, it grows another on the opposite side after molting. [23] A soft abdomen is also present in a hermit crab as an asymmetrical modification due to the asymmetrical nature of the snail shells they inhabit. [24]
All gastropods are asymmetrical. This is easily seen in snails and sea snails, which have helical shells. At first glance slugs appear externally symmetrical, but their pneumostome (breathing hole) is always on the right side. The origin of asymmetry in gastropods is a subject of scientific debate. [25] Other gastropods develop external asymmetry, such as Glaucus atlanticus that develops asymmetrical cerata as they mature.
Histioteuthis is a genus of squid, commonly known as the cock-eyed squid, because in all species the right eye is normal-sized, round, blue and sunken; whereas the left eye is at least twice the diameter of the right eye, tubular, yellow-green, faces upward, and bulges out of the head.
Corals build colonies that are not symmetrical, but the individual polyps exhibit radial symmetry. [26]
Alpheidae feature asymmetrical claws that lack pincers, the larger of which can grow on either side of the body, and if lost can develop on the opposite arm instead. [27]
Certain polyopisthocotylean monogeneans are asymmetrical, as an adaptation to their attachment to the gill of their fish hosts.[ citation needed ]
Certain parasitic copepods [ which? ] which live inside the gill chamber of their fish hosts are asymmetrical.[ citation needed ]
Lobsters of the genera Homarus , Nephrops , and Homarinus have dimorphic claws, a crushing one and a cutting one. [28]
Thrips have asymmetrical mouthparts, unique among insects. [29]
Invertebrates is an umbrella term describing animals that neither develop nor retain a vertebral column, which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordate subphylum Vertebrata, i.e. vertebrates. Well-known phyla of invertebrates include arthropods, mollusks, annelids, echinoderms, flatworms, cnidarians and sponges.
Cichlids are fish from the family Cichlidae in the order Cichliformes. Traditionally Cichlids were classed in a suborder, the Labroidei, along with the wrasses (Labridae), in the order Perciformes, but molecular studies have contradicted this grouping. On the basis of fossil evidence, it first appeared in Tanzania during the Eocene epoch, about 46–45 million years ago. The closest living relative of cichlids is probably the convict blenny, and both families are classified in the 5th edition of Fishes of the World as the two families in the Cichliformes, part of the subseries Ovalentaria. This family is large, diverse, and widely dispersed. At least 1,650 species have been scientifically described, making it one of the largest vertebrate families. New species are discovered annually, and many species remain undescribed. The actual number of species is therefore unknown, with estimates varying between 2,000 and 3,000.
Lake Tanganyika is an African Great Lake. It is the second-oldest freshwater lake in the world, the second-largest by volume, and the second-deepest, in all cases after Lake Baikal in Siberia. It is the world's longest freshwater lake. The lake is shared among four countries—Tanzania, the Democratic Republic of the Congo (DRC), Burundi, and Zambia, with Tanzania (46%) and DRC (40%) possessing the majority of the lake. It drains into the Congo River system and ultimately into the Atlantic Ocean.
Asymmetry is the absence of, or a violation of, symmetry. Symmetry is an important property of both physical and abstract systems and it may be displayed in precise terms or in more aesthetic terms. The absence of or violation of symmetry that are either expected or desired can have important consequences for a system.
Sinistral and dextral, in some scientific fields, are the two types of chirality ("handedness") or relative direction. The terms are derived from the Latin words for "left" (sinister) and "right" (dexter). Other disciplines use different terms or simply use left and right.
Symmetry in biology refers to the symmetry observed in organisms, including plants, animals, fungi, and bacteria. External symmetry can be easily seen by just looking at an organism. For example, the face of a human being has a plane of symmetry down its centre, or a pine cone displays a clear symmetrical spiral pattern. Internal features can also show symmetry, for example the tubes in the human body which are cylindrical and have several planes of symmetry.
The Florida stone crab is a crab found in the western North Atlantic, from Connecticut to Colombia, including Texas, the Gulf of Mexico, Belize, Mexico Jamaica, Cuba, The Bahamas, and the East Coast of the United States. The crab can also be found in and around the salt marshes of South Carolina and Georgia. It is widely caught for food. The closely related species Menippe adina is sometimes considered a subspecies – they can interbreed, forming hybrids – and they are treated as one species for commercial fishing, with their ranges partly overlapping. The two species are believed to have diverged approximately 3 million years ago.
A molluscivore is a carnivorous animal that specialises in feeding on molluscs such as gastropods, bivalves, brachiopods and cephalopods. Known molluscivores include numerous predatory molluscs,, arthropods such as crabs and firefly larvae, and, vertebrates such as fish, birds and mammals. Molluscivory is performed in a variety ways with some animals highly adapted to this method of feeding behaviour. A similar behaviour, durophagy, describes the feeding of animals that consume hard-shelled or exoskeleton bearing organisms, such as corals, shelled molluscs, or crabs.
Lepidophagy is a specialised feeding behaviour in fish that involves eating the scales of other fish. Lepidophagy is widespread, having evolved independently in at least five freshwater families and seven marine families. A related feeding behavior among fish is pterygophagy: feeding on the fins of other fish.
Iwasaki's snail-eater is a species of snake in the family Pareidae. The species is endemic to the Yaeyama Islands in the southern Ryukyu Islands, Japan.
Perissodus microlepis is a species of cichlid endemic to Lake Tanganyika. This species reaches a length of 11 centimetres (4.3 in) TL. This species can also be found in the aquarium trade. It is a scale-eating 'parasite' on other fish species. It occurs in two distinct morphological forms. One morph has mouth parts twisted to the left, enabling it to eat scales off its victim's right flank. In contrast, the other morph, whose mouth is twisted to the right, eats scales off its victim's left flank. The relative abundance of the two morphs in populations is regulated by frequency-dependent selection.
Aetiocetus is a genus of extinct basal mysticete, or baleen whale that lived 33.9 to 23.03 million years ago, in the Oligocene in the North Pacific ocean, around Japan, Mexico, and Oregon, U.S. It was first described by Douglas Emlong in 1966 and currently contains known four species, A. cotylalveus, A. polydentatus, A. tomitai, and A. weltoni. These whales are remarkable for their retention of teeth and presence of nutrient foramina, indicating that they possessed baleen. Thus, Aetiocetus represents the transition from teeth to baleen in Oligocene mysticetes. Baleen is a highly derived character, or synapomorphy, of mysticetes, and is a keratinous structure that grows from the palate, or roof of the mouth, of the whale. The presence of baleen is inferred from the fossil record in the skull of Aetiocetus. Aetiocetus is known from both sides of the Pacific Ocean: it was first documented in Oregon, United States, but it is also known from Japan and Mexico. The genus is currently constrained to the Northern hemisphere and has little value in biostratigraphic studies of the Oligocene due to its limited occurrences across the Pacific.
Durophagy is the eating behavior of animals that consume hard-shelled or exoskeleton bearing organisms, such as corals, shelled mollusks, or crabs. It is mostly used to describe fish, but is also used when describing reptiles, including fossil turtles, placodonts and invertebrates, as well as "bone-crushing" mammalian carnivores such as hyenas. Durophagy requires special adaptions, such as blunt, strong teeth and a heavy jaw. Bite force is necessary to overcome the physical constraints of consuming more durable prey and gain a competitive advantage over other organisms by gaining access to more diverse or exclusive food resources earlier in life. Those with greater bite forces require less time to consume certain prey items as a greater bite force can increase the net rate of energy intake when foraging and enhance fitness in durophagous species.
The Mammalia in the 10th edition of Systema Naturae forms one of six classes of animals in Carl Linnaeus's tenth reformed edition written in Latin. The following explanations are based on William Turton's translations who rearranged and corrected earlier editions published by Johann Friedrich Gmelin, Johan Christian Fabricius and Carl Ludwig Willdenow:
Animals that suckle their young by means of lactiferous teats. In external and internal structure they resemble man: most of them are quadrupeds; and with man, their natural enemy, inhabit the surface of the Earth. The largest, though fewest in number, inhabit the ocean.
Chirality is a property of asymmetry important in several branches of science. The word chirality is derived from the Greek χειρ (kheir), "hand", a familiar chiral object.
Most bony fishes have two sets of jaws made mainly of bone. The primary oral jaws open and close the mouth, and a second set of pharyngeal jaws are positioned at the back of the throat. The oral jaws are used to capture and manipulate prey by biting and crushing. The pharyngeal jaws, so-called because they are positioned within the pharynx, are used to further process the food and move it from the mouth to the stomach.
Pareidae is a small family of snakes found largely in southeast Asia, with an isolated subfamily endemic to southwestern India. It encompasses 42 species in four genera divided into two subfamilies: Pareinae and Xylophiinae. Both families are thought to have diverged from one another during the early-mid Eocene, about 40-50 million years ago.
The contralateral organization of the forebrain is the property that the hemispheres of the cerebrum and the thalamus represent mainly the contralateral side of the body. Consequently, the left side of the forebrain mostly represents the right side of the body, and the right side of the brain primarily represents the left side of the body. The contralateral organization involves both executive and sensory functions. The contralateral organization is only present in vertebrates.
Left-right asymmetry is the process in early embryonic development that breaks the normal symmetry in the bilateral embryo. In vertebrates, left-right asymmetry is established early in development at a structure called the left-right organizer and leads to activation of different signalling pathways on the left and right of the embryo. This in turn cause several organs in adults to develop LR asymmetry, such as the tilt of the heart, the different number lung lobes on each side of the body and the position of the stomach and spleen on the right side of the body. If this process does not occur correctly in humans it can result in the syndromes heterotaxy or situs inversus.