Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of Eumillipes persephone, which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures.
Pauropods are small, pale, millipede-like arthropods. Around 830 species in twelve families are found worldwide, living in soil and leaf mold. They look rather like centipedes, but are probably the sister group to millipedes. However, this is controversial, as a close relationship with Symphyla has also been posited.
Glomerida is an order of pill-millipedes found primarily in the Northern Hemisphere. They superficially resemble pill-bugs or woodlice, and can enroll into a protective ball. They have twelve body segments, 17 to 19 pairs of legs, and males have enlarged rear legs involved in mating. The order includes about 30 genera and at least 280 species, including Glomeris marginata, the common European pill-millipede. The order contains members in Europe, South-east Asia and the Americas from California to Guatemala. Although historically considered closely related with the similar sphaerotheriidans that also enroll, some DNA evidence suggest they may be more closely related to glomeridesmidans, a poorly known order that does not enroll.
Polydesmida is the largest order of millipedes, containing approximately 3,500 species, including all the millipedes reported to produce hydrogen cyanide (HCN).
Glomeridesmida is an order of millipedes in the infraclass Pentazonia containing 2 families and at least 31 species. Glomeridesmida is the only living order of the superorder Limacomorpha. Glomeridesmidans are small and somewhat flattened, and unlike other orders of Pentazonia, are unable to roll into a ball. Ocelli (eyes) are absent.
Chordeumatida is a large order of millipedes containing some 1200 species with a nearly worldwide distribution. Also known as "sausage millipedes," they possess around 30 body segments behind the head as adults and reach about 25 mm (0.98 in) in length.
Siphoniulus is a poorly known genus of millipede containing only two living species: S. alba from Indonesia, and S. neotropicus from Mexico and Guatemala. An additional two fossil species are known from Cretaceous amber. Siphoniulus species are the only members of the family Siphoniulidae and order Siphoniulida, making Siphoniulida the smallest millipede order. Few specimens are known, and their classification is contentious, although most recent studies place them as basal members of the Helminthomorpha.
Lankasomatidae, is a millipede family in the suborder Heterochordeumatidea of order Chordeumatida. Adult millipedes in this family have 32 segments, not the 30 segments usually found in this order. The family includes 11 species belongs to three genera.
Craspedosomatidae is a family of millipedes in the order Chordeumatida. Most adult millipedes in this family have 30 segments, but some have only 28. There are at least 30 genera and 210 described species in Craspedosomatidae.
Trichopetalidae is a family of millipedes in the order Chordeumatida. Adult millipedes in this family have 28 or 30 segments. There are about 5 genera and at least 30 described species in Trichopetalidae.
Metopidiotrichidae is a family of millipedes in the order Chordeumatida. Adult millipedes in this family have 32 segments, not the 30 segments usually found in this order. Adult males in this family often feature a reduced or vestigial leg pair 10 as part of the gonopod complex, in addition to the two leg pairs typically modified into gonopods in this order. There are about 9 genera and at least 70 described species in Metopidiotrichidae.
Haasiidae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have 28 or 30 segments.
Apterouridae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have 30 segments.
Pygmaeosomatidae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have 30 or 32 segments.
Adritylidae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have 30 segments.
Hoffmaneumatidae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have only 28 segments, not the 30 segments usually found in this order. Adult males in this family feature a reduced leg pair 10 in addition to the two pairs normally modified into gonopods in this order. In the adult male of the species Hoffmaneuma exiguum, for example, the gonopod complex derives from all three leg pairs rather than from just the usual two.
Biokoviellidae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have 30 segments.
Heterolatzeliidae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have 30 segments.
Opisthocheiridae is a family of millipedes belonging to the order Chordeumatida. Adult millipedes in this family have 26 or 30 segments. This family includes the species Opisthocheiron canayerensis, one of the few species in this order with only 26 segments.
