| Sphaerotherium giganteum | |
|---|---|
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Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Subphylum: | Myriapoda |
| Class: | Diplopoda |
| Order: | Sphaerotheriida |
| Family: | Sphaerotheriidae |
| Genus: | Sphaerotherium |
| Species: | S. giganteum |
| Binomial name | |
| Sphaerotherium giganteum Porat, 1872 | |
Sphaerotherium giganteum (the giant pill millipede) is a species of southern African pill millipede in the family Sphaerotheriidae. [1] [2]
The giant pill millipede is the largest species of pill millipede in South Africa. Its length is 5 cm, with the body segmented into 11-13 segments. Each segment has two pairs of legs, for a total of about 42 legs. [1]
The giant pill millipede is found in moist forest habitat under leaf litter in South Africa. It is widely distributed in high-altitude forested areas in Kwa-Zulu Natal, Limpopo, Mpumalanga and the Eastern Cape. [1]
The giant pill millipede feeds on decaying matter on the forest floor. The carnivorous slug, Chlamydephorus , is known to prey on S. giganteum in the coastal forests of KZN. [1]
The population number of Sphaerotherium giganteum has not been determined, and it is not formally protected. Sphaerotherium giganteum may be vulnerable to habitat loss and degradation, as well as exploitation in the pet trade and in traditional medicine. [1]
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from Latin for "thousand feet", no species was known to have 1,000 or more until the discovery in 2020 of Eumillipes persephone, which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures.
Centipedes are predatory arthropods belonging to the class Chilopoda of the subphylum Myriapoda, an arthropod group which includes millipedes and other multi-legged animals. Centipedes are elongated segmented (metameric) creatures with one pair of legs per body segment. All centipedes are venomous and can inflict painful stings, injecting their venom through pincer-like appendages known as forcipules or toxicognaths, which are actually modified legs instead of fangs. Despite the name, no species of centipede has exactly 100 legs; the number of pairs of legs is an odd number that ranges from 15 pairs to 191 pairs.
Myriapods are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial.
Pill millipedes are any members of two living orders of millipedes, often grouped together into a single superorder, Oniscomorpha. The name Oniscomorpha refers to the millipedes' resemblance to certain woodlice (Oniscidea), also called pillbugs or "roly-polies". However, millipedes and woodlice are not closely related ; rather, this is a case of convergent evolution.
Arthropleura is a genus of massive myriapod that lived in what is now Europe and North America around 345 to 290 million years ago, from the Viséan stage of the lower Carboniferous Period to the Sakmarian stage of the lower Permian Period. It is related to millipedes, and was capable of reaching at least 2 metres in length, possibly up to over 2.5 metres, making it the largest known land arthropod of all time. Arthropleura is known from body fossils as well as trace fossils, particularly giant trackways up to 50 centimetres (20 in) wide, and potentially also large burrows. It lived in open, sparsely wooded environments near water, and was possibly amphibious.
Sphaerotheriidae is a family of giant pill millipedes of the class Diplopoda. Millipedes of this family are distributed in southern Africa.
Glomeris marginata is a common European species of pill millipede. It is a short millipede, rounded in cross-section, which is capable of rolling itself up into a ball ("volvation") when disturbed. This behaviour is also found in the pill woodlouse Armadillidium, with which G. marginata is often confused.
Archispirostreptus gigas, known as the giant African millipede, shongololo or Bongololo, is the largest extant species of millipede, growing up to 33.5 centimetres (13.2 in) in length, 67 millimetres (2.6 in) in circumference. It has approximately 256 legs, although the number of legs changes with each molting so it can vary according to each individual.
Glomerida is an order of pill-millipedes found primarily in the Northern Hemisphere. Also known as northern pill millipedes, they superficially resemble pill-bugs or woodlice, and can enroll into a protective ball. They have twelve body segments, 17 to 19 pairs of legs, and males have enlarged rear legs involved in mating. The order includes about 30 genera and at least 280 species, including Glomeris marginata, the common European pill-millipede. The order contains members in Europe, South-east Asia and the Americas from California to Guatemala. Although historically considered closely related with the similar sphaerotheriidans that also enroll, some DNA evidence suggest they may be more closely related to glomeridesmidans, a poorly known order that does not enroll.
Chlamydephorus is a genus of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the family Chlamydephoridae. It is the only genus within the family Chlamydephoridae.
Sphaerotheriida is an order of millipedes in the infraclass Pentazonia, sometimes known as giant pill millipedes. They inhabit Southern Africa, Madagascar, South and Southeast Asia, Australia and New Zealand. Like the Northern Hemisphere pill millipedes of the order Glomerida, these millipedes can roll into a ball when disturbed. When they are rolled-up, most sphaerotheriidans reach a maximum size of a cherry or golf ball, but some species from Madagascar can even reach the size of an orange. When rolled-up, predators are unable to unravel giant pill millipedes since the margins of their second and last dorsal plates fit perfectly into one another, creating a sealed ball. A few giant pill millipede species are able to produce sound, the only millipedes known to do this. This order of millipedes is also unique in that some African species are used for medicinal purposes.
Sphaeromimus, or the chirping giant pill millipedes, is a genus of giant pill millipedes endemic to southeastern Madagascar. Though described in 1902, the genus was up to 2005 known from a single male specimen, whose appearance was so unusual that the authors suspected a mislabeled giant pill-millipede from India. Their unusual and distinct morphology includes well-developed stridulation organs, probably as devices for courtship. These are the male ‘harp’ and the female ‘washboard’, which contain more stridulation ribs than in other members of the order Sphaerotheriida. They have a closer affinity with the Indian genus Arthrosphaera than to other Malagasy genera, though all belong to the Arthrosphaeridae.
Crurifarcimen is a monotypic genus of millipedes containing the single species Crurifarcimen vagans. Its common name is wandering leg sausage. This millipede is endemic to Tanzania, where it is known only from the Usambara Mountains. It was formally described in 2011 and placed in a newly erected genus of its own. It was declared one of the world's top 10 new species of 2012 by Arizona State University's International Institute for Species Exploration.
Polyzoniida is an order of millipedes in the subterclass Colobognatha. This order contains three families and more than 70 described species. The species in this order are also known as camphor millipedes, because ozopore secretions in this order frequently have a strong camphor-like smell. Poison frogs in South America and Madagascar evidently obtain some of their poison from these millipedes.
Arthrosphaeridae is a family of giant pill millipedes that are found in Madagascar, the Western Ghats of India, and Sri Lanka. The largest pill-millipede in the world, Zoosphaerium neptunus belongs to this family and is known to swarm at certain times of the year.
Zoosphaerium neptunus, also known as the giant emerald pill millipede, is a species of millipede within the family Arthrosphaeridae. Its size is an example of island gigantism, it being the largest known pill-millipede in the world, with some individuals reaching a maximum length of 90 mm long. The species is endemic to Madagascar, where it is known to swarm at certain times of the year. Z. neptunus is the only known species within the order Sphaerotheriida that is known to exhibit this mass swarming behaviour.
Zoosphaerium darthvaderi, also known as the Darth Vader giant pill millipede, is a species of giant pill millipede endemic to the island state of Madagascar. It is named after the Star Wars character Darth Vader. The millipede is restricted to the Ambohitantely Reserve and is considered an extremely endangered species.
Glomeris sublimbata, also known as the marbled pill millipede, is a species of pill millipede belonging to the genus Glomeris and the family Glomeridae.
Zoosphaerium album, also known as the giant opal pill millipede, is a species of pill millipede within the family Arthrosphaeridae and order Sphaerotheriida. It is an endemic species, found only within the Vohibasia forest of Western Madagascar. The species name album, refers to the white colouration of the pill millipedes exoskeleton.
Telodeinopus autii, commonly known as the giant African olive millipede, the Ghana speckled leg millipede, and the long legged millipede, is a species of large millipede inhabiting the tropical forests of west, central and east Africa.