Zygodactylidae | |
---|---|
Primozygodactylus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Aves |
Clade: | Psittacopasseres |
Family: | † Zygodactylidae Brodkorb, 1971 |
Genera | |
†Eozygodactylus | |
Synonyms | |
Primoscenidae Mayr 1998 |
Zygodactylidae is a family of extinct birds found in Europe and North America from the Eocene epoch to the Middle Miocene. [1] First named in 1971, based on fragmentary remains of two species from Germany, a more complete description of the birds became possible in 2008 when a number of other, better-preserved fossil species were assigned to the family based on a number of shared characteristics. [2]
The name of the group comes from the presence of a zygodactyl foot, with two toes projecting forward, and two to the rear. This is the same arrangement as seen in living parrots and woodpeckers, and the zygodactylids were at one time thought to be related to the woodpecker family. More recent analyses, however, have shown that they are more likely to be the sister clade to the passerine or "perching" birds, the large clade that includes, among others, all living songbirds. [3] Passerine birds are distinguished by an anisodactyl foot, but it is thought that their earliest ancestors may have been zygodactyl, likely using this arrangement of toes to cling to the bark of trees as woodpeckers do. Only later did the fourth toe rotate from the rear to face forward in the modern passerine fashion. [4]
† Family Zygodactylidae
A passerine is any bird of the order Passeriformes which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their toes, which facilitates perching.
The turacos make up the bird family Musophagidae ( "banana-eaters"), which includes plantain-eaters and go-away-birds. In southern Africa both turacos and go-away-birds are commonly known as loeries. They are semi-zygodactylous: the fourth (outer) toe can be switched back and forth. The second and third toes, which always point forward, are conjoined in some species. Musophagids often have prominent crests and long tails; the turacos are noted for peculiar and unique pigments giving them their bright green and red feathers.
The mousebirds are birds in the order Coliiformes. They are the sister group to the clade Cavitaves, which includes the Leptosomiformes, Trogoniformes (trogons), Bucerotiformes, Piciformes and Coraciformes. This group is now confined to sub-Saharan Africa, and it is the only bird order confined entirely to that continent, with the possible exception of turacos which are considered by some as the distinct order Musophagiformes, and the cuckoo roller, which is the only member of the order Leptosomiformes, and which is found in Madagascar but not mainland Africa. Mousebirds had a wider range in the Paleogene, with a widespread distribution in Europe and North America during the Paleocene.
Lithornithidae is an extinct, possibly paraphyletic group of early paleognath birds. They are known from fossils dating to the Upper Paleocene through the Middle Eocene of North America and Europe, with possible Late Cretaceous representatives. All are extinct today; the youngest specimen is the currently unnamed SGPIMH MEV1 specimen from the mid-Eocene Messel Pit site.
Psittacopes is an extinct genus of bird from Middle Eocene. One species is recorded from Messel, Germany, and other three possible species are from London Clay, England, one named ?Psittacopes occidentalis in 2022, and the other two unnamed. Its phylogenetic placement within Aves is uncertain; it was originally interpreted as a parrot, but the phylogenetic analysis conducted by Mayr (2015) recovered it as more closely related to the passerines and the extinct family Zygodactylidae.
Eurofluvioviridavis is a genus of extinct primitive birds from the Middle Eocene Messel Pit, Germany. It contains a single species, Eurofluvioviridavis robustipes. It is related to Avolatavis and Vastanavis, other members of the family Vastanavidae.
Messelasturidae is an extinct family of birds known from the Eocene of North America and Europe. Their morphology is a mosaic that in some aspects are very similar to modern hawks and falcons, but in others are more similar to parrots. Initially interpreted as stem-owls, more recent studies have suggested a closer relationship to parrots and passerines. Their ecology is enigmatic.
Vastanavidae is an extinct family of birds related to parrots and passerine birds. They are known from fossils from Eocene sites in India, Europe, and North America. The vastanavids resemble parrots and the extinct parrot relative Quercypsitta in their morphology, including the partially zygodactyl foot, in which two toes could face opposite the other two.
Halcyornithidae is an extinct family of telluravian birds thought to be related to the Psittaciformes (parrots), Passeriformes (songbirds), and to the extinct Messelasturidae. Halcyornithids have been found in various Eocene formations in Europe and North America. Widespread and diverse in the Early Eocene of North America and Europe, halcyornithids are not found in locales later than the Middle Eocene. Halcyornithids were small, arboreal birds with zygodactyl feet, with two toes facing forwards and two facing back, a trait shared with other tree-dwelling families of Eocene birds like the Zygodactylidae and the messelasturids. The skull of halcyornithids features a ridge of bone above the eye called the supraorbital process, similar to birds of prey. The relationships of the halcyornithids to other birds remain uncertain. Halcyornithids have been proposed as relatives to owls and as a lineage closer to parrots than to songbirds. Most recently, halcyornithids have been identified as the sister group of the clade including parrots and songbirds. It is also possible that Halcyornithidae is paraphyletic with respect to the Messelasturidae.
Geranoididae is a clade of extinct birds from the early to late Eocene and possibly early Oligocene of North America and Europe. These were mid-sized, long-legged flightless birds. Recent research shows that these birds may actually be palaeognaths related to ostriches.
Morsoravis is an extinct genus of neoavian bird from the Early Eocene Fur Formation of Denmark. It contains a single named species, Morsoravis sedilis. Fossils of Morsoravis have also been found in the Green River Formation of Wyoming and possibly the Nanjemoy Formation of Virginia.
This is an overview of the paleofauna of the Eocene Messel Formation as explored by the Messel Pit excavations in Germany. A former quarry and now UNESCO World Heritage Site, the Messel Formation preserves what once were a series of anoxic lakes surrounded by a sub-tropical rainforest during the Middle Eocene, approximately 47 Ma.
Minutornis is an extinct genus of parapasserine bird from the Early Eocene London Clay Formation of Essex, United Kingdom. The genus contains a single species, M. primoscenoides, known from a fragmentary skeleton.
Cyrilavis is an extinct genus of halcyornithid bird from the Early Eocene Fossil Butte Member of the Green River Formation, United States. The genus contains two species, Cyrilavis olsoni and Cyrilavis colburnorum.
Pseudasturides is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany, and possibly the Isle of Sheppey, United Kingdom. The genus is represented by a single species, Pseudasturides macrocephalus.
Serudaptus is an extinct genus of halcyornithid bird from the Middle Eocene Messel pit in Hesse, Germany. The genus contains one species, Serudaptus pohli, and is known for long, raptorial claws on its zygodactyl feet.
Parapsittacopes is an extinct genus of psittacopedid bird from the Early Eocene London Clay, United Kingdom. The genus contains one species, Parapsittacopes bergdahli.
Eofringillirostrum is an extinct species of bird known from the Early Eocene Green River Formation of the Western United States and from the Messel Pit in Germany. The genus contains two species, Eofringillirostrum boudreauxi and Eofringillirostrum parvulum. They are the earliest known fossil birds to have a finch-like beak.
Sororavis is an extinct genus of bird from the Early Eocene London Clay of Walton-on-the-Naze, United Kingdom. It contains a single named species, Sororavis solitarius. It is a member of the Morsoravidae.
Morsoravidae is a family of extinct birds known from the Early Eocene of Europe and North America, related to the Psittacopasseres, the clade containing parrots and songbirds. They have specialised feeding ecologies, and one morsoravid, Pumiliornis tesselatus, fed on nectar. The family contains the genera Morsoravis, Pumiliornis, and Sororavis.