| Lessemsaurus Temporal range: Norian ~ | |
|---|---|
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Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Clade: | Dinosauria |
| Clade: | Saurischia |
| Clade: | † Sauropodomorpha |
| Clade: | † Sauropoda |
| Clade: | † Lessemsauridae |
| Genus: | † Lessemsaurus Bonaparte 1999 |
| Species: | †L. sauropoides |
| Binomial name | |
| †Lessemsaurus sauropoides Bonaparte 1999 | |
Lessemsaurus is an extinct genus of sauropodiform dinosaur belonging to Lessemsauridae.
The type species, L. sauropoides, was formally described by José Fernando Bonaparte in 1999 in honor of Don Lessem, a writer of popular science books. It was found in the Los Colorados Formation of the Ischigualasto-Villa Unión Basin in La Rioja Province, Argentina. [1]
It was discovered in strata dating to the Norian stage, around 210 million years ago. [2] [3] It is estimated to have reached 10–12 metres (33–39 ft) long and weighed over 7 metric tons (7.7 short tons), possibly up to 8–10 metric tons (8.8–11.0 short tons), in maximum body mass. [4] [5] [6] [7]
A cladogram after Pol, Garrido & Cerda, 2011, [8] illustrates a possible placing of Lessemsaurus and Antetonitrus in Sauropodomorpha:
In 2018, Apaldetti et al. recovered it as part of a clade they named Lessemsauridae, after Lessemsaurus. Their cladogram is reproduced below: [9]
The following cladogram shows the position of Lessemsaurus outside of Sauropoda, according to Oliver W. M. Rauhut and colleagues, 2020: [10]
Eoraptor is a genus of small, lightly built, basal sauropodomorph dinosaur. One of the earliest-known dinosaurs and one of the earliest members of the sauropod family, it lived approximately 231 to 228 million years ago, during the Late Triassic in Western Gondwana, in the region that is now northwestern Argentina. The type and only species, Eoraptor lunensis, was first described in 1993, and is known from an almost complete and well-preserved skeleton and several fragmentary ones. Eoraptor had multiple tooth shapes, which suggests that it was omnivorous. Eoraptor was 1.5 feet (0.46 m) tall and 3 feet (0.91 m) long.
Sauropodomorpha is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had long necks and tails, were quadrupedal, and became the largest animals to ever walk the Earth. The prosauropods, which preceded the sauropods, were smaller and were often able to walk on two legs. The sauropodomorphs were the dominant terrestrial herbivores throughout much of the Mesozoic Era, from their origins in the Late Triassic until their decline and extinction at the end of the Cretaceous.
Saturnalia is an extinct genus of basal sauropodomorph dinosaur known from the Late Triassic Santa Maria Formation of Rio Grande do Sul, southern Brazil. It is one of the earliest known dinosaurs.
Melanorosaurus is a genus of basal sauropodomorph dinosaur that lived during the Late Triassic period. A herbivore from South Africa, it had a large body and sturdy limbs, suggesting it moved about on all fours. Its limb bones were massive and heavy like the limb bones of true sauropods.
Guaibasaurus is an extinct genus of basal saurischian dinosaur known from the Late Triassic Caturrita Formation of Rio Grande do Sul, southern Brazil. Most analyses recover it as a sauropodomorph, although there are some suggestions that it was a theropod instead. In 2016 Gregory S. Paul estimated it at 2 meters and 10 kg, whereas in 2020 Molina-Pérez and Larramendi listed it at 3 meters and 35 kg.
Antetonitrus is a genus of sauropodiform dinosaur found in the Early Jurassic Elliot Formation of South Africa. The only species is Antetonitrus ingenipes. Sometimes considered a basal sauropod, it is crucial for the understanding of the origin and early evolution of this group. It was a quadrupedal herbivore, like its later relatives, but shows primitive adaptations to use the forelimbs for grasping, instead of purely for weight support.
Blikanasaurus is a genus of sauropodomorph dinosaur from the late Triassic of South Africa. The generic name Blikanasaurus is derived from Greek, meaning "lizard from Blikana". The species name cromptoni is taken from the surname of A.W. "Fuzz" Crompton, an American paleontologist who led numerous field expeditions in Elliot Formation outcrop localities in South Africa. Blikanasaurus is only known from partial hindlimb bones that were recovered from the lower Elliot Formation (LEF) in the Eastern Cape.
Coloradisaurus is a genus of massospondylid sauropodomorph dinosaur. It lived during the Late Triassic period in what is now La Rioja Province, Argentina. It is known from two specimens collected from the Los Colorados Formation of the Ischigualasto-Villa Unión Basin.
Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and Antarctica during the Late Triassic to the Early Jurassic periods. Several dinosaurs have been classified as massospondylids over the years. The largest cladistic analysis of early sauropodomorphs, which was presented by Apaldetti and colleagues in November 2011, found Adeopapposaurus, Coloradisaurus, Glacialisaurus, Massospondylus, Leyesaurus and Lufengosaurus to be massospondylids. This result supports many previous analyses that tested fewer taxa. However, this analysis found the two recently described North American massopods, Sarahsaurus and Seitaad, and the South African Ignavusaurus to nest outside Massospondylidae, as opposed to some provisional proposals. Earlier in 2011, Pradhania, a sauropodomorph from India, was tested for the first time in a large cladistic analysis and was found to be a relatively basal massospondylid. Mussaurus and Xixiposaurus may also be included within Massospondylidae.
Anchisauria is an extinct clade of sauropodomorph dinosaurs that lived from the Late Triassic to the Late Cretaceous. The name Anchisauria was first used Haekel and defined by Galton and Upchurch in the second edition of The Dinosauria. It is a node-based taxon containing the most recent common ancestor of Anchisaurus polyzelus and Melanorosaurus readi, and all its descendants. Galton and Upchurch assigned a family of dinosaurs to the Anchisauria: the Melanorosauridae. The more common prosauropods Plateosaurus and Massospondylus were placed in the sister clade Plateosauria.
Plateosauria is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to the Late Cretaceous. The name Plateosauria was first coined by Gustav Tornier in 1913. The name afterwards fell out of use until the 1980s.
Massopoda is a clade of sauropodomorph dinosaurs which lived during the Late Triassic to Late Cretaceous epochs. It was named by paleontologist Adam M. Yates of the University of the Witwatersrand in 2007. Massopoda is a stem-based taxon, defined as all animals more closely related to Saltasaurus loricatus than to Plateosaurus engelhardti.
The Quebrada del Barro Formation is a geological formation of the Marayes-El Carrizal Basin in San Juan Province, Argentina. This formation is the most fossiliferous portion of the Triassic Marayes Group, and is also the youngest unit of the group, overlying the El Carrizal Formation. An unconformity at the top of the Quebrada del Barro Formation separates it from the Cretaceous-age Los Riscos Formation of the El Gigante Group. Part of the formation may be made into a provincial park following the discovery of the fossils of Ingentia, a giant sauropodomorph dinosaur which helped elucidate the early evolution of sauropods.
Leonerasaurus is a basal genus of sauropodomorph dinosaur. Currently, there is only one species known, named L. taquetrensis by Diego Pol, Alberto Garrido and Ignacio A. Cerda in 2011. The fossil, an incomplete subadult individual, was found in the Las Leoneras Formation in Argentina. This formation is probably Early Jurassic in age. Leonerasaurus was a small non-sauropod sauropodomorph, showing an unusual combination of basal and derived characters. This indicates that the evolution of early sauropodomorphs witnessed a great degree of convergent evolution.
Leyesaurus is an extinct genus of massospondylid sauropodomorph dinosaur known from the San Juan Province, northwestern Argentina.
Xingxiulong is a genus of bipedal sauropodiform from the Early Jurassic of China. It contains a single species, X. chengi, described by Wang et al. in 2017 from three specimens, two adults and an immature individual, that collectively constitute a mostly complete skeleton. Adults of the genus measured 4–5 metres (13–16 ft) long and 1–1.5 metres tall. Phylogenetic analysis suggests that Xingxiulong is most closely related to its contemporary Jingshanosaurus, although an alternative position outside of both the Sauropodiformes and Massospondylidae is also plausible.
Ingentia is a genus of early sauropodiform dinosaur, sometimes considered a basal sauropod, from the Late Triassic of Argentina. The type specimen of Ingentia, PVSJ 1086, was discovered in the Quebrada del Barro Formation of northwestern Argentina. It was described in 2018 by Cecilia Apaldetti, Ricardo Nestor Martínez, Ignacio Alejandro Cerda, Diego Pol and Oscar Alcober who named the type and only species Ingentia prima, meaning "first huge one", as the taxon was one of the first very large sauropodomorphs to evolve, along with its close relative Lessemsaurus. A second specimen, PVSJ 1087, was referred, containing five tail vertebrae, both ulnae and radii, a left calfbone and a right foot.
Lessemsauridae is a clade of early sauropodiform dinosaurs that lived in the Triassic and Jurassic of Argentina, South Africa and possibly Lesotho. A phylogenetic analysis performed by Apaldetti and colleagues in 2018 recovered a new clade of sauropodiforms uniting Lessemsaurus, Antetonitrus, and Ingentia which they named Lessemsauridae. It is a node-based taxon, defined as all descendants of the most recent common ancestor of Lessemsaurus sauropoides and Antetonitrus ingenipes. Depending on the definition of Sauropoda, Lessemsauridae is either one of the most basal sauropod taxa, or a sister taxon of Sauropoda. An additional member of the clade was named later in 2018, Ledumahadi. A 2021 study by Pol and colleagues also assigned the genera Kholumolumo and Meroktenos to the group.
Ledumahadi is a genus of lessemsaurid sauropodomorph dinosaur from the Early Jurassic Elliot Formation in Free State Province, South Africa. The type and only species is L. mafube, known from a singular incomplete postcranial specimen. A quadruped, it was one of the first giant sauropodomorphs, reaching a weight of around 12 tonnes, despite not having evolved columnar limbs like its later huge relatives.
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