Shingopana

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Shingopana
Temporal range: late Campanian-early Maastrichtian
~74.3–71.4  Ma
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Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha
Clade: Sauropoda
Clade: Macronaria
Clade: Titanosauria
Clade: Lithostrotia
Clade: Aeolosaurini
Genus: Shingopana
Gorscak et al. 2017
Type species
Shingopana songwensis
Gorscak et al. 2017

Shingopana (meaning "wide neck" in Swahili) is a genus of titanosaurian sauropod from the Upper Cretaceous (late Campanian-early Maastrichtian [2] age) Galula Formation of Tanzania. It is known from only the type species, S. songwensis. [3] Gorscak & O'Connor's phylogenetic testing suggest Shingopana is more closely related to the South American titanosaur family of Aeolosaurini than any of the titanosaurs found so far in North & South Africa. [3]

Discovery and naming

Part of the holotype, TZ-07, was discovered in 2002 by scientists affiliated with the Rukwa Rift Basin Project, which was run by Patrick O'Connor and Nancy Stevens. [3] The rest of the skeleton was excavated during the following years. The species Shingopana songwensis was officially named in 2017. [3]

Description

Shingopana was a quadrupedal Aeolosaurin sauropod that would have reached up to 8 metres (26 ft) long when fully grown, smaller than the average sauropod. [3]

Skeleton

The holotype was damaged by insect bore holes shortly after the animal died. [3]

Shingopana is known from a partial jaw, represented by the angular bone. Shingopana is also known from four cervical vertebrae; with two of these vertebrae having preserved cervical ribs and another isolated cervical rib. Shingopana instead had remnants of a bulbous expansion on the incompletely preserved cervical vertebrae, which probably helped to strengthen its neck. [3]

Four ribs have been preserved with the holotype, but none are complete. The ribs had flanged edges, but their function is currently unknown. [3]

An almost complete humerus and a partial pubis were also present in the holotype.

Classification

Palaeoecology

The holotype was discovered in the Upper Cretaceous Galula Formation of the Rukwa Rift Basin in Tanzania. It would have coexisted with the sauropods Rukwatitan [4] and Mnyamawamtuka, [5] the mesoeucrocodiles Pakasuchus [6] and Rukwasuchus , [7] the mammal Galulatherium , [4] an unnamed notosuchian, an unnamed turtle, an unnamed theropod [4] and two types of lungfish ( Lupaceradotus and an unnamed genus).

See also

Related Research Articles

<i>Alamosaurus</i> Extinct genus of dinosaurs

Alamosaurus is a genus of opisthocoelicaudiine titanosaurian sauropod dinosaurs containing a single known species, Alamosaurus sanjuanensis, from the Maastrichtian age of the Late Cretaceous period in what is now southwestern North America. Isolated vertebrae and limb bones indicate that it reached sizes comparable to Argentinosaurus and Puertasaurus, which would make it the absolute largest dinosaur known from North America. Its fossils have been recovered from a variety of rock formations spanning the Maastrichtian age. Specimens of a juvenile Alamosaurus sanjuanensis have been recovered from only a few meters below the Cretaceous-Paleogene boundary in Texas, making it among the last surviving non-avian dinosaur species. Alamosaurus is the only known sauropod to have inhabited North America after their nearly 30-million year absence from the North American fossil record and probably represents an immigrant from South America.

<i>Euhelopus</i> Extinct genus of dinosaurs

Euhelopus is a genus of sauropod dinosaur that lived between 145 and 133 million years ago during the Berriasian and Valanginian stages of the Early Cretaceous in what is now Shandong Province in China. It was a large quadrupedal herbivore. Like sauropods such as brachiosaurs and titanosaurs, Euhelopus had longer forelegs than hind legs. This discovery was paleontologically significant because it represented the first dinosaur scientifically investigated from China: seen in 1913, rediscovered in 1922, and excavated in 1923 and studied by T'an during the same year. Unlike most sauropod specimens, it has a relatively complete skull.

<span class="mw-page-title-main">Lithostrotia</span> Extinct clade of dinosaurs

Lithostrotia is a clade of derived titanosaur sauropods that lived during the Early Cretaceous and Late Cretaceous. The group was defined by Upchurch et al. in 2004 as the most recent common ancestor of Malawisaurus and Saltasaurus and all the descendants of that ancestor. Lithostrotia is derived from the Ancient Greek lithostros, meaning "inlaid with stones", referring to the fact that many known lithostrotians are preserved with osteoderms. However, osteoderms are not a distinguishing feature of the group, as the two noted by Unchurch et al. include caudal vertebrae with strongly concave front faces (procoely), although the farthest vertebrae are not procoelous.

<i>Galulatherium</i> Fossil taxon

Galulatherium is an extinct genus of possibly gondwanathere mammal, from the Late Cretaceous (Turonian-Campanian)-aged Galula Formation of Tanzania. It is known solely from the type specimen TNM 02067 a fragmentary fossil dentary. The short, deep bone is about 19.5 mm (0.77 in) long, but the back part is broken off. It contains a large, forward-inclined incisor with a root that extends deep into the jaw, separated by a diastema (gap) from five cheekteeth. Very little remains of the teeth, but enough to determine that they are hypsodont (high-crowned). The third cheektooth is the largest and the roots of the teeth are curved. First described in 2003, TNM 02067 has been tentatively identified as a sudamericid—an extinct family of high-crowned gondwanathere mammals otherwise known from South America, Madagascar, India, and Antarctica. If truly a gondwanathere, it would be the only African member of the group and may be the oldest. The describers could not exclude other possibilities, such as that the jaw represents some mammalian group known only from younger, Cenozoic times. In 2019 the fossil was CT scanned, which revealed additional details of the specimen.

<i>Pakasuchus</i> Extinct genus of reptiles

Pakasuchus is a genus of notosuchian crocodyliform distinguished by its unusual mammal-like appearance, including mammal-like teeth that would have given the animal the ability to chew. It also had long, slender legs and a doglike nose. Fossils have been found in the Galula Formation of Rukwa Rift Basin of southwestern Tanzania, and were described in 2010 in the journal Nature. Pakasuchus is originally considered to lived approximately 105 million years ago, in the mid-Cretaceous, but later age of site is reconsidered to the late Cretaceous, Cenomanian to Campanian instead. The type species is P. kapilimai. Pakasuchus means "cat crocodile" in reference to its catlike skull.

<i>Angolatitan</i> Extinct genus of titanosauriform dinosaurs

Angolatitan is a genus of titanosauriform sauropod dinosaur from the Upper Cretaceous. It is also the first non-avian dinosaur discovered in Angola. The genus contains a single species, Angolatitan adamastor, known from a partial right forelimb. Angolatitan was a relict form of its time; it was a Late Cretaceous basal titanosauriform, when more derived titanosaurs were far more common.

<span class="mw-page-title-main">Aeolosaurini</span> Extinct clade of dinosaurs

Aeolosaurini is an extinct clade of titanosaurian dinosaurs known from the Cretaceous period of Argentina and Brazil. Rodrigo M. Santucci and Antonio C. de Arruda-Campos (2011) in their cladistic analysis found Aeolosaurus, Gondwanatitan, Maxakalisaurus, Panamericansaurus and Rinconsaurus to be aeolosaurids.

Tataouinea is an extinct genus of sauropod dinosaur in the subfamily Rebbachisaurinae of Rebbachisauridae which lived in the Early Cretaceous of Tunisia. Only one species, T. hannibalis, is known.

Rukwasuchus is an extinct genus of peirosaurid mesoeucrocodylian known from the middle Cretaceous Galula Formation of southwestern Tanzania. It contains a single species, Rukwasuchus yajabalijekundu.

<i>Rukwatitan</i> Extinct genus of dinosaurs

Rukwatitan is a genus of titanosaur sauropod dinosaur from the Galula Formation in Tanzania. It lived around 100 million years ago, during the middle Cretaceous. The species, which shared features with another southern African species, Malawisaurus dixeyi, measured 30 feet (9.1 m) from the head to the tip of the tail, and had forelimbs that were estimated around 6.5 feet (2.0 m) long. Its fossils were found embedded in a cliff face near Lake Rukwa in the Rukwa Valley, from which it gets its name.

Lohuecotitan is an extinct genus of titanosaurian sauropod dinosaur which lived during the Late Cretaceous in Spain. The only species known in the genus is Lohuecotitan pandafilandi, described and named in 2016.

<i>Mansourasaurus</i> Extinct genus of dinosaurs

Mansourasaurus is a genus of herbivorous lithostrotian sauropod dinosaur from the Quseir Formation of Egypt. The type and only species is Mansourasaurus shahinae.

The Quseir Formation is a geological formation in the vicinity of the Kharga Oasis in Egypt. It is Campanian In age. The lithology largely consists of soft shale with hard bands of sandstone, siltstone and phosphorite. The environment of deposition was nearshore to freshwater fluvio-lacustrine characterized by moist and aquatic habitats with a tropical warm-humid climate. It is conformably overlain by the marine late Campanian-Maastrichtian Duwi Formation, and unconformably overlies the Turonian Taref Formation. The sauropod dinosaurs Mansourasaurus and Igai are known from the formation, as well as the proximal fibula of an indeterminate theropod. Additionally the lungfish genera Lavocatodus and Protopterus, the crocodyliform Wahasuchus and the bothremydid turtle Khargachelys are also known.

The Galula Formation is a geological formation located south of Lake Rukwa in Tanzania, part of the Red Sandstone Group of the Rukwa Rift Basin. Along with the unconformably overlying Oligocene Nsungwe Formation. It is divided into two members, the lower Mtuka Member and the upper Namba Member.

Mnyamawamtuka is a genus of lithostrotian titanosaur sauropod dinosaur from the Cretaceous Galula Formation in Tanzania. The type and only species is M. moyowamkia.

The Nsungwe Formation is a formation in the Rukwa Rift Basin of the East African Rift System, it is Oligocene in age based on U-Pb dating of a tuff horizon within the formation. It is part of the Red Sandstone Group along with the uncomfortably underlying Mid-Cretaceous Galula Formation It is divided into two members, the lower Utengule Member, and the upper Songwe member. It is notable for being one of the most important Paleogene fossil deposits in Sub-Saharan Africa.

<i>Abditosaurus</i> Genus of titanosaurian dinosaur

Abditosaurus is an extinct genus of titanosaurian sauropod dinosaur from the Late Cretaceous (Maastrichtian) Tremp Group of Catalonia, Spain. The type and only species is Abditosaurus kuehnei. Phylogenetic analyses recover it within a clade of South American and African saltasaurines, distinct from other insular dwarf sauropods from the European archipelago. Abditosaurus inhabited the Ibero-Armorican Island, a prehistoric island made up of what is now Spain, Portugal, and southern France, and would have been the largest titanosaur species in its environment.

Ruixinia is an extinct genus of somphospondylan titanosauriform dinosaur from the Early Cretaceous (Barremian) Yixian Formation of China. The genus contains a single species, Ruixinia zhangi. The Ruixinia holotype is a partial articulated skeleton with the most complete series of caudal vertebrae known from any Asian titanosauriform.

<i>Jiangxititan</i> Genus of somphospondylan sauropod dinosaurs

Jiangxititan is an extinct genus of somphospondylan titanosauriform dinosaur from the Late Cretaceous Nanxiong Formation of China. The genus contains a single species, J. ganzhouensis, known from several articulated vertebrae with ribs. Originally described as a titanosaur, Jiangxititan was later suggested to be a non-titanosaurian somphospondylan.

<i>Inawentu</i> Genus of titanosaurian dinosaurs

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References

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  2. Widlansky, Sarah J.; Clyde, William C.; O'Connor, Patrick M.; Roberts, Eric M.; Stevens, Nancy J. (2018-03-01). "Paleomagnetism of the Cretaceous Galula Formation and implications for vertebrate evolution". Journal of African Earth Sciences. 139: 403–420. Bibcode:2018JAfES.139..403W. doi: 10.1016/j.jafrearsci.2017.11.029 . ISSN   1464-343X.
  3. 1 2 3 4 5 6 7 8 Gorscak, E.; O'Connor, P. M.; Roberts, E. M.; Stevens, N. J. (2017). "The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania, with remarks on African titanosaurian diversity". Journal of Vertebrate Paleontology. 361 (4): 35–55. Bibcode:2017JVPal..37E3250G. doi:10.1080/02724634.2017.1343250. S2CID   90885040.
  4. 1 2 3 O’Connor, Patrick M.; Gottfried, Michael D.; Stevens, Nancy J.; Roberts, Eric M.; Ngasala, Sifa; Kapilima, Saidi; Chami, Remigius (March 2006). "A new vertebrate fauna from the Cretaceous Red Sandstone Group, Rukwa Rift Basin, Southwestern Tanzania". Journal of African Earth Sciences. 44 (3): 277–288. Bibcode:2006JAfES..44..277O. doi:10.1016/j.jafrearsci.2005.11.022. ISSN   1464-343X.
  5. Gorscak, Eric; O’Connor, Patrick M. (2019). "A new African titanosaurian sauropod dinosaur from the middle Cretaceous Galula Formation (Mtuka Member), Rukwa Rift Basin, southwestern Tanzania". PLoS ONE . 2 (14): e0211412. Bibcode:2019PLoSO..1411412G. doi: 10.1371/journal.pone.0211412 . PMC   6374010 . PMID   30759122.
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  7. Sertich, Joseph J. W.; O’Connor, Patrick M. (2014-04-16). "A new crocodyliform from the middle Cretaceous Galula Formation, southwestern Tanzania". Journal of Vertebrate Paleontology. 34 (3): 576–596. Bibcode:2014JVPal..34..576S. doi:10.1080/02724634.2013.819808. ISSN   0272-4634. S2CID   16644660.