Alsophila javanica

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Alsophila javanica
Scientific classification OOjs UI icon edit-ltr.svg
Kingdom: Plantae
Clade: Tracheophytes
Division: Polypodiophyta
Class: Polypodiopsida
Order: Cyatheales
Family: Cyatheaceae
Genus: Alsophila
Species:
A. javanica
Binomial name
Alsophila javanica
Synonyms [1]
  • Alsophila benculensisAlderw.
  • Cyathea javanicaBlume
  • Alsophila palembanicaAlderw.
  • Cyathea barisanica(Alderw.) Domin
  • Cyathea benculensisAlderw.
  • Cyathea caudipinnula(Alderw.) Domin
  • Cyathea palembanicaAlderw.
  • Hemitelia barisanicaAlderw.
  • Hemitelia caudipinnulaAlderw.

Alsophila javanica, synonym Cyathea javanica, [1] is a species of tree fern native to western Java and Sumatra, in Indonesia. It grows in rain forest and on riverbanks at an altitude of 250–1500 m.

Description

The trunk of Alsophila javanica is erect and up to about 10 m tall. Fronds may be bi- or tripinnate and 2–3 m in length. The stipe is spiny and bears scattered scales throughout. These scales are dark and have fragile edges. A. javanica has round sori which are borne near the midvein of fertile pinnules. They are protected by firm, flat indusia that are saucer-like in appearance. [2]

Taxonomy

Alsophila javanica appears to be most closely related to Alsophila doctersii . Large and Braggins (2004) note that A. doctersii may in fact be of hybrid origin between A. javanica and a member of the Alsophila latebrosa complex. [2]

When Carl Blume described A. javanica (as Cyathea javanica), he also named one variety, rigida. Large and Braggins (2004) note that although variety rigida is apparently part of the A. javanica complex, the name has also occasionally been associated with Alsophila polycarpa . [2]

Related Research Articles

<i>Alsophila australis</i> Species of fern

Alsophila australis, synonym Cyathea australis, also known as the rough tree fern, is a species of tree fern native to southeastern Queensland, New South Wales and southern Victoria in Australia, as well as Tasmania and Norfolk Island.

<i>Alsophila cunninghamii</i> Species of fern

Alsophila cunninghamii, synonym Cyathea cunninghamii, also known as the gully tree fern and slender tree fern, is a species of tree fern indigenous to New Zealand including North Island, South Island and Chatham Islands; also to Victoria, possibly New South Wales, southeastern Queensland and Tasmania in Australia. It grows in damp forest, often emerging from stream gullies and riverbanks. Brownsey noted that it has a lower tolerance for drought than other related species. The erect trunk may be 20 m tall and is usually 6–15 cm in diameter, occasionally as much as 20 cm. Fronds are tri- to tetrapinnate and 3 m or more in length. The rachis and stipe are slender, black brown, warty and covered with brown scales. Sori occur along each side of the pinnule midvein and are covered by hood-like indusia. A. cunninghamii is an uncommon and slow-growing tree fern.

Alsophila doctersii, synonym Cyathea doctersii, is a species of tree fern endemic to Sumatra, where it grows in forest at an altitude of approximately 150 m. The trunk is erect and 2–3 m tall. Fronds are bi- or tripinnate and 2–3 m long. The stipe is covered with scattered flat, brown scales and some hairs. Sori occur near the fertile pinnule midvein and are covered by thin, brown indusia that are scale-like in appearance.

Alsophila dryopteroides, synonym Cyathea dryopteroides, is a tree fern native to Puerto Rico, where it grows in shaded areas and cloud forests at an altitude of 1000–1200 m. The erect trunk may be 1.3 m tall and approximately 5 cm in diameter. Fronds are pinnate and up to 1.6 m long. The rachis is often purplish brown and covered with scales, usually on the underside. The scales range in colour from golden brown to bicoloured. Sori occur along each side of the pinnule midvein and indusia are cup-like.

Alsophila fulgens, synonym Cyathea fulgens, is a species of tree fern endemic to the Dominican Republic on the island of Hispaniola. It forms part of the complex centered on Alsophila woodwardioides comprising six very similar taxa from the Greater Antilles. The other five species are Alsophila jimeneziana, Alsophila grevilleana, Alsophila portoricensis and Alsophila tussacii. Large and Braggins (2004) note that this group is known to cross with members of the Alsophila minor complex. In the wild, A. fulgens also forms hybrids with Alsophila brooksii.

Alsophila geluensis, synonym Cyathea geluensis, is a species of tree fern native to central and eastern New Guinea as well as the Louisiade Archipelago, where it grows in mossy forest. In New Guinea itself, plants grow at an altitude of 1000–2000 m, however they are present at lower elevations of 700–900 m on associated islands. The trunk of this tree fern is erect and may be 5 m tall or more. Fronds are bi- or tripinnate and 1–2.5 m long. They are usually about ten live fronds present in the crown at once. The stipe may be warty and/or have short spines as well as many scattered scales towards the base. These scales are pale to dark and have dull, fragile edges. Sori occur near the fertile pinnule midvein and are protected by pale, thin indusia. A. geluensis is a variable taxon and further study is needed to determine whether it does not in fact represent a species complex.

Gymnosphaera gigantea, synonyms Alsophila gigantea and Cyathea gigantea, is a species of tree fern native to northeastern to southern India, Sri Lanka, Nepal to Myanmar, Thailand, Laos, Vietnam, the Malay Peninsula, as well as central Sumatra and western Java. It grows in moist open areas at an elevation of 600–1000 m. The trunk of this species is erect and may be as tall as 5 m or more. Fronds are bi- or tripinnate and usually 2–3 m long. The rachis is long, dark to black in colouration and rough in appearance after the fall of scales. These scales are dark brown, glossy and have a narrow paler margin and fragile edges. Sori are round and indusia absent.

Gymnosphaera glabra, synonyms Alsophila glabra and Cyathea glabra, is a species of tree fern native to Borneo, western Java, Sumatra, the Lesser Sunda Islands, Sulawesi, and the Malay Peninsula, where it grows in lowland swamp forest and montane forest at an elevation of up to 1500 m. The trunk of this plant is erect and 2–4 m tall. Fronds are bi- or tripinnate and 1–2 m in length. Characteristically of this species, the lowest pinnae may be significantly reduced. The stipe is very dark and bears basal scales. These scales are dark, glossy and have a paler margin and fragile edges. Sori are produced in groups of one to three on fertile pinnule veins. They lack indusia.

Alsophila gleichenioides, synonym Cyathea gleichenioides, is a species of tree fern endemic to New Guinea, where it grows in open peaty grassland and on forest margins, often in groups, at an altitude of 2800–3700 m. The trunk of this plant is erect, up to 3 m tall and about 24 cm in diameter. The narrow fronds are tripinnate and about 1 m in length. Around 60 fronds form a rounded crown. The stipe is warty and bears scattered scales towards the base. These scales may be either glossy brown with a paler dull margin, or small, pale and fringed. Sori occur one or two per fertile pinnule and are protected by firm, brown indusia.

Alsophila grevilleana, synonym Cyathea grevilleana, is a species of tree fern endemic to Jamaica, where it grows in moist gullies and on wooded hills in both calcareous and noncalcareous soils at an altitude of 200–1200 m. The trunk of this plant is erect, about 7 m tall, and 10–15 cm in diameter. It is characteristically clothed in old stipe bases, brown scales and blackish spines. Fronds are tripinnate, dark green in colour, and up to 4 m in length. The last pinnae are sometimes separated, forming a distinctive clump around the trunk apex. The rachis is yellow-brown and almost smooth. This species has a long, dark brown stipe with a few scattered spines. Sori are produced in four to six pairs along the pinnule midvein. They are protected by pale brown indusia that are cup-like in appearance.

Alsophila heterochlamydea, synonym Cyathea heterochlamydea, is a little-known species of tree fern native to the islands of Luzon, Panay, Negros and Mindanao in the Philippines, where it grows in montane forest. The trunk of this plant is erect and usually up to 4 m tall or more. Fronds may be bi- or tripinnate and 1–2 m in length. The stipe is warty and/or bears short spines and scales. These scales are dark, glossy and have a narrow pale margin. Sori are borne near the fertile pinnule midvein and are protected by firm, brown indusia.

Alsophila × marcescens, synonym Cyathea × marcescens, commonly known as the skirted tree fern, is a tree fern endemic to the Cape Otway ranges in Victoria and Tasmania, Australia. It is a natural hybrid, apparently Alsophila australis × Alsophila cunninghamii. Large and Braggins (2004) note that it has characteristics midway between these two species. The spores of A. × marcescens are usually malformed although sterile. The trunk of this plant is erect and up to 10 m tall. Fronds may be bi- or tripinnate and 3–4 m in length. Dead fronds often persist, forming a characteristic skirt around the trunk. The stipe is thick, black and warty. The rachis and trunk are covered in shiny, dark brown scales. Sori are borne near the fertile pinnule midvein and are protected by thin indusia that are saucer-like in appearance.

Alsophila humilis, synonym Cyathea humilis, is a species of tree fern native to Kenya, as well as the Usambara and Uluguru Mountains in Tanzania, where it grows in wet forest at an altitude of 1100–2000 m. The trunk of this plant is erect and 2–3 m tall. Fronds are pinnate and 1–2 m in length. Dead fronds are often retained in the typical variety, forming an irregular skirt around the trunk. The rachis and stipe are light brown in colouration. Scales are present towards the base of the stipe. They are dark, glossy, and have narrow, fragile edges. Sori occur at the forks of veins and are protected by thin, reduced indusia.

Alsophila brausei, synonym Cyathea hunsteiniana, is a species of tree fern endemic to eastern New Guinea, where it grows in rain forest at an altitude of 1300–2000 m. This species has a slender, erect trunk up to 1 m tall and about 3 cm in diameter. Fronds are bipinnate and approximately 1 m in length. Scattered scales cover the stipe. They are dark brown in colouration and have a broad paler margin and fragile edges. Round sori are borne one or two per fertile pinnule segment. The sori are protected by deep, firm indusia that are cup-like in appearance.

Alsophila imbricata, synonym Cyathea imbricata, is a species of tree fern endemic to Western New Guinea, where it grows in open forest at an altitude of 3240 m. The trunk of this plant is erect and approximately 2 m tall. Fronds may be bi- or tripinnate and are usually less than 1 m in length. The stipe is dark, spiny, and covered with caducous scales. These scales are glossy brown in colouration and have a paler margin and fragile edges. Sori are borne in groups of one to four per pinnule lobe. They are protected by firm indusia.

<i>Alsophila incisoserrata</i> Species of fern

Alsophila incisoserrata, synonym Cyathea incisoserrata, is a species of tree fern native to the Malay Peninsula and the state of Sarawak on the island of Borneo, where it grows in forest and forest margins from the lowland to approximately 1250 m. The trunk is erect, about 4 m tall and 12 cm in diameter. Fronds may be bi- or tripinnate and 1–2 m in length. The stipe of this species is partly persistent and is sometimes retained on the upper trunk. It is warty, bears conical spines, and is sparsely covered with scales. These scales may be either small and fringed or bullate. Sori are borne near the fertile pinnule midvein. They are protected by very small, often bilobed indusia.

Alsophila junghuhniana, synonym Cyathea junghuhniana, is a species of tree fern native to southern and central Sumatra and western Java, where it grows in forest at an altitude of 1000–2000 m. The trunk of this species is erect and may be 2 m tall or more. Fronds are tripinnate and up to about 3 m long. The stipes of this species are persistent and form a skirt around the trunk. They are brown, spiny, and covered with glossy, dark brown scales. Sori are borne near the midvein of fertile pinnules and are protected by thin indusia.

Alsophila arfakensis, synonym Cyathea kanehirae, is a species of tree fern native to western New Guinea, where it grows in montane forest at an altitude of 1600–2700 m. The trunk of this plant is erect and 1–4 m tall. Fronds may be bi- or tripinnate and 1–2 m in length. The rachis is smooth, while the stipe is dark and warty. The stipe is covered with flat, brown, scattered scales. Sori are borne near the fertile pinnule midvein. They are protected by small indusia that are cup-like in appearance.

<i>Alsophila latebrosa</i> Species of fern

Alsophila latebrosa, synonym Cyathea latebrosa, is a common and widespread species of tree fern native to Indochina. Its natural range covers Cambodia and Thailand, and stretches from the Malay Peninsula to Indonesia, where it is present on the islands of Borneo and Sumatra. Plants reported from India and Sri Lanka have thinner indusia and may represent a separate, as-yet undescribed, species. A. latebrosa grows in a wide range of habitats, including forest, secondary forest, and plantations, from sea level up to an elevation of about 1500 m.

Alsophila crassicaula, synonym Cyathea ledermannii, is a species of tree fern native to Papua New Guinea and Bougainville Province in the Solomon Islands, where it is common in submontane rain forest at an altitude of 1000–3000 m. The trunk of this plant is erect and grows to about 3 m in height. Fronds may be bi- or tripinnate and up to 2 m in length. The rachis is purplish brown in colouration and usually bears basal scales. These scales range from pale, to brown, to bicoloured. Sori are borne on each side of the pinnule midvein. They are protected by firm indusia.

References

  1. 1 2 3 Hassler, Michael & Schmitt, Bernd (June 2019). "Alsophila javanica". Checklist of Ferns and Lycophytes of the World. Vol. 8. Archived from the original on 2017-09-02. Retrieved 2019-08-26.
  2. 1 2 3 Large, Mark F. & Braggins, John E. (2004). Tree Ferns. Timber Press. p.  149. ISBN   978-0-88192-630-9.