Armenoceratidae

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Armenoceratidae
Temporal range: M Ordovician - Silurian
Orthoceras cochleatus.jpg
Part of the isolated siphuncle of an armenoceratid nautiloid cephalopod
Scientific classification Red Pencil Icon.png
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Nautiloidea
Order: Actinocerida
Family: Armenoceratidae
Troedsson (1926)
Genera

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The Armenoceratidae are a family of early Paleozoic nautiloid cephalopods belonging to the order Actinocerida. [1] [2]

The Armenoceratidae, established by Troedsson (1926) [2] are characterized by large, straight, or slightly curved shells and large siphuncles with strongly expanded segments between the septa. Septal necks are short and abruptly recurved along brims. Radial canals in the endosiphuncular canal system are typically arched, curving forward and backward from near the septal foramina (openings) to connect with the parispatium on either side of the middle of each segments. [2] The parispatium is the narrow opening between the inner side of the connecting rings in actinocerids and the internal siphuncular deposits that grow forward and back from the region of the septal openings.

The Armenoceratidae have their beginning in Armenoceras which first appeared near the beginning of the Chazyan, 2nd stage of the Middle Ordovician in older established chronologies, in northeastern China, Manchuria, [1] [3] derived from Wutinoceras . Additional genera include Nybyoceras and Selkirkoceras , respectively, from the upper Middle and Upper Ordovician, and Monocyrtoceras , Elrodoceras , and Megadiscoceras from the Silurian, [2]

The Armenoceratidae, Armenoceras , Nybyoceras , and Selkirkoceras , first appear in North America in the diverse cephalopod Redriveran faunas of the early Upper Ordovician [1] [3] but are no longer found during that stage in Asia. Selkirkoceras is a large armenocerid from the Upper Ordovician with a blunt, flattened, somewhat breviconic shell.

Among the Silurian genera, Elrodoceras has a large shell with the apical part slightly curved, otherwise is straight, and a siphuncle that is narrower than in Armenoceras. Monocyrtoceras has a siphuncle like that of Elrodoceras, but the entire shell is gently and evenly curved. Megadiscosorus is similar to Armenoceras, but more breviconic and slightly exogastrically cyrtoconic and with the siphuncle in contact with the ventral wall.

The Armenoceratidae most likely gave rise through Armenoceras to Gonioceras in the Chazyan, and later in the Middle Ordovician, possibly through either an early Armenoceras or Nybyoceras to Actinoceras

Related Research Articles

Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.

Actinoceras is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician. It is an extinct genus of nautiloid cephalopod that thrived in the warm waters of the United States during the Paleozoic era.

<i>Ormoceras</i> Extinct genus of molluscs

Ormoceras is an actinocerid nautiloid genus and type for the family Ormoceratidae, found in North America from the late Chazyan through the early Cincinnatian of the Middle and Upper Ordovician, but which continued through the Devonian worldwide.

Lambeoceras is a genus of large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America and the sole representative of the family Lambeoceratidae.

Armenoceras is a genus of actinocerid nautiloid cephalopods whose fossils ranged from the late Whiterockian Stage in the early Middle Ordovician, through the remainder of the period and on into the Upper Silurian. It is the type genus of the family Armenoceratidae.

Oncocerida Extinct order of nautiloids

The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian, in which the connecting rings are thin and siphuncle segments are variably expanded. At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three.

Bathmoceras is a primitive cephalopod genus from the Middle and Upper Ordovician, type for the family Bathmoceratidae, order Ellesmerocerida.

The Actinocerida are an order of generally straight, medium to large cephalopods that lived during the early and middle Paleozoic, distinguished by a siphuncle composed of expanded segments that extend into the adjacent chambers, in which deposits formed within contain a system of radial canals and a narrow space along the inner side of the connecting ring known as a paraspatium. Septal necks are generally short and cyrtochoanitic, some being recumbent, some hook shaped. Most grew to lengths of about 60 to 90 cm but some, like the Huroniidae of the Silurian grew significantly larger.

Ellesmeroceratidae Extinct family of molluscs

The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic).

The Reudemannoceratidae are the ancestral and most primitive of the Discosorida, an order of cephalopods from the early Paleozoic. The Reudemannoceratidae produced generally medium-sized endogastric and almost straight shells with the siphuncle slightly ventral from the center.

Oncoceratidae Extinct family of nautiloids

Oncoceratidae is a family of nauatiloid cephalopods in the order Oncocerida established by Hyatt, 1884, that range from the Middle Ordovician to the Upper Silurian.

The Actinoceriatidae are a family of actinocerids named by Saemann in 1853 for those that grew to have large shells with blunt apices and large siphuncles with widely expanded segments and a generally arcuate endosiphucular canal system. Their range is from the upper Middle Ordovician to the Lower Silurian. Actinocerids are generally straight-shelled nautiloid cephalopods with a siphuncle composed of expanded segments, typically with thin connecting rings, in which the internal deposits are penetrated by a system of canals

Nybyoceras is an actinocerid genus assigned to the Armenoceratidae and similar to Armenoceras except for having a siphuncle close to the ventral side of the shell.

Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.

Mandaloceratidae is a family in the nautiloid cephalopod order Discosorida, from the Middle and Upper(?) Silurian characterized by short, essentially straight shells referred to as breviconic, typically with a faintly exogastric shape produced by the profile of the body chamber.

Ascoceratidae is a family of Ascocerida, bizarre orthoceratoid cephalopods, with longiconic, deciduous early growth stages which undergo period truncation and an inflated breviconic mature ascoceroid stage. The Ascoceratidae are divided into two, unequal subfamilies.

Cyrtogomphoceras is a genus of nautiloid cephalopods, recognized by its large breviconic shell with a notable endogastric curvature. The shell is fusiform in profile, reaching maximum width at or near the base of body chamber, which narrows toward the aperture. The siphuncle is large and slightly removed from the ventral side, that with the concave longitudinal profile. Siphuncle segments are short, as are chambers; septal necks recurved, connecting rings thick, bullettes at the apical end of the rings swollen. Cameral deposits are lacking.

Elrodoceras is a genus of armenoceratids,, that grew fairly large, characterized by a straight shell, slightly curved at the apex, a siphuncle narrower than that of Armenoceras but with segments still wider than long, and an arched endosiphuncular canal system.

Monocyrtoceras is a genus of armenoceratids (Cephalopoda) from the Middle Silurian of North America (Wisconsin), with a gently curved shell and a siphuncle like that of Elrodoceras.

Megadisocosorus is a genus of actinocerid cephalopods similar to Armenoceras but with a short, slightly curved, breviconic shell. The siphuncle is in contact with the ventral wall, unlike that of Armenoceras in which the siphuncle is close to the center.

References

  1. 1 2 3 Flower 1957.Studies of the Actinoceratida.; Memoir 2; New Mexico Bureau of Mines and Mineral Resources, Socorro, NM
  2. 1 2 3 4 Teichert 1964. Actinoceratoidea, Treatise on Invertebrate Paleontology, part K.(Nautiloidea)
  3. 1 2 Flower 1976 Ordovician Cephalopod Faunas and Their Role in Correlation. in Basset M. C. (ed) The Ordovician System. Palaeonological Association.