Atlas wild ass | |
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Atlas wild asses in a Roman mosaic of El Jem, shown attacked by a tiger | |
Extinct (ca. 300 AD) | |
Scientific classification ![]() | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Perissodactyla |
Family: | Equidae |
Genus: | Equus |
Species: | |
Subspecies: | †E. a. atlanticus |
Trinomial name | |
†Equus africanus atlanticus P. Thomas, 1884 |
The Atlas wild ass (Equus africanus atlanticus), also known as Algerian wild ass, is a purported extinct subspecies of the African wild ass that was once found across North Africa and parts of the Sahara. [1]
It was last represented in a villa mural ca. 300 AD in Bona, Algeria, and may have become extinct as a result of Roman sport hunting. [2]
Purported bones have been found in a number of rock shelters across Morocco and Algeria by paleontologists including Alfred Romer (1928, 1935) and Camille Arambourg (1931). [3] While the existence of numerous prehistoric rock art depictions, and Roman mosaics leave no doubt about the former existence of African wild asses in North Africa, it has been claimed that the original bones that were used to describe the subspecies atlanticus actually belonged to a fossil zebra. Therefore, the name E. a. atlanticus would be "unavailable" to the Atlas wild ass. [1] [4]
It was also hypothesized that the appearance of Nubian and Somali wild asses were clinal and that they appeared different as an artifact of the recent extinction of intermediate-looking populations. This would make the living African wild ass a monotypic species with no subspecies, and at least question the existence of extinct subspecies like the Atlas wild ass. However, genetic studies have shown since that Nubian and Somali wild asses are different enough to warrant subspecies status. Additionally, domestic donkeys carry two different haplotypes, one shared with the Nubian wild ass, and another of unknown origin that is not found in the Somali wild ass. The presence of the extinct Atlas wild ass in the Ancient Mediterranean makes it a plausible source for the second haplotype. [4] [5] [6]
Ancient art consistently depicts the African wild asses of North Africa as similar to, but darker colored than, the Nubian and Somali wild ass subspecies. The general color was gray, with marked black and white stripes on the legs, and a black shoulder cross [7] (sometimes doubled). [4] In comparison, the Nubian wild ass is gray with shoulder cross but no stripes, and the Somali wild ass is sandy with black stripes, but no shoulder cross. [1] One or both features appear occasionally in domestic donkeys. Wild and primitive domestic asses are indistinguishable from their bones, which complicates their identification in archaeological sites. [4]
The Atlas wild ass was found in the region around the Atlas Mountains, across modern day Algeria, Tunisia and Morocco. [8] It might also have occurred in rocky areas of the Saharan Desert, but not in sands which are avoided by wild asses. [1] However, the 20th century reports of wild asses from northern Chad and the Hoggar Massif in the central Sahara are doubtful. [4]
Equidae is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. The family evolved around 50 million years ago from a small, multi-toed ungulate into larger, single-toed animals. All extant species are in the genus Equus, which originated in North America. Equidae belongs to the order Perissodactyla, which includes the extant tapirs and rhinoceros, and several extinct families.
The quagga is an extinct subspecies of the plains zebra that was endemic to South Africa until it was hunted to extinction in the late 19th century. It was long thought to be a distinct species, but early genetic studies have supported it being a subspecies of plains zebra. A more recent study suggested that it was the southernmost cline or ecotype of the species.
Zebras are African equines with distinctive black-and-white striped coats. There are three living species: Grévy's zebra, the plains zebra, and the mountain zebra. Zebras share the genus Equus with horses and asses, the three groups being the only living members of the family Equidae. Zebra stripes come in different patterns, unique to each individual. Several theories have been proposed for the function of these patterns, with most evidence supporting them as a deterrent for biting flies. Zebras inhabit eastern and southern Africa and can be found in a variety of habitats such as savannahs, grasslands, woodlands, shrublands, and mountainous areas.
The donkey or ass is a domesticated equine. It derives from the African wild ass, Equus africanus, and may be classified either as a subspecies thereof, Equus africanus asinus, or as a separate species, Equus asinus. It was domesticated in Africa some 5000–7000 years ago, and has been used mainly as a working animal since that time.
The onager, also known as hemione or Asiatic wild ass, is a species of the family Equidae native to Asia. A member of the subgenus Asinus, the onager was described and given its binomial name by German zoologist Peter Simon Pallas in 1775. Six subspecies have been recognized, two of which are extinct.
The plains zebra is the most common and geographically widespread species of zebra. Its range is fragmented, but spans much of southern and eastern Africa south of the Sahara. Six or seven subspecies have been recognised, including the extinct quagga which was thought to be a separate species. More recent research supports variations in zebra populations being clines rather than subspecies.
Equus is a genus of mammals in the family Equidae, which includes horses, asses, and zebras. Within the Equidae, Equus is the only recognized extant genus, comprising seven living species. Like Equidae more broadly, Equus has numerous extinct species known only from fossils. The genus originated in North America and dispersed into the Old World and South America during the Early and Middle Pleistocene. Equines are odd-toed ungulates with slender legs, long heads, relatively long necks, manes, and long tails. All species are herbivorous, and mostly grazers, with simpler digestive systems than ruminants but able to subsist on lower-quality vegetation.
The maned rat or (African) crested rat is a nocturnal, long-haired and bushy-tailed East African rodent that superficially resembles a porcupine. The world's only known poisonous rodent, the maned rat sequesters toxins from plants to fend off predators.
The red gazelle is an extinct species or subspecies of gazelle. It was formerly considered a member of the genus Gazella within the subgenus Eudorcas before Eudorcas was promoted to a full genus. It is thought to have lived in the better-watered mountain areas of North Africa rather than in deserts, because of the rich colouring on the coat.
The wild asses (Asinus) are a subgenus of single toed grazing ungulates. Its species are:
The African wild ass or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey, which is sometimes placed within the same species. They live in the deserts and other arid areas of the Horn of Africa, in Eritrea, Ethiopia and Somalia. It formerly had a wider range north and west into Sudan, Egypt, and Libya. It is Critically Endangered, with about 570 existing in the wild.
The Somali wild ass is a subspecies of the African wild ass.
Asinus is a subgenus of Equus that encompasses several subspecies of the Equidae commonly known as wild asses, characterized by long ears, a lean, straight-backed build, lack of a true withers, a coarse mane and tail, and a reputation for considerable toughness and endurance.
The Syrian wild ass, less commonly known as a hemippe, an achdari, or a Mesopotamian or Syrian onager, is an extinct subspecies of onager native to the Arabian peninsula and surrounding areas. It ranged across present-day Iraq, Palestine, Israel, Iran, Jordan, Saudi Arabia, Syria, and Turkey.
The bubal hartebeest, also known as northern hartebeest or bubal antelope or simply bubal is the extinct nominal subspecies of hartebeest. It was formerly found north of the Saharan Desert. Other subspecies live currently in grasslands south of the Sahara, from Senegal in the west to Eritrea and Ethiopia in the east and down to central Tanzania. The red hartebeest and Lichtenstein's hartebeest, alternatively considered subspecies or sister species of the common hartebeest, are present in southern Africa.
The Nubian wild ass is the nominate subspecies of African wild ass, and one of the ancestors of the domestic donkey, which was domesticated about 6,000 years ago. It is presumed to be extinct, though two populations potentially survive on the Caribbean island of Bonaire and in Gebel Elba.
Primitive markings are a group of hair coat markings and qualities seen in several equine species, including horses, donkeys, and asses. In horses, they are associated with primitive breeds, though not limited to such breeds. The markings are particularly associated with the dun coat color family. All dun horses possess at least the dorsal stripe, but the presence of the other primitive markings varies. Other common markings may include horizontal striping on the legs, transverse striping across the shoulders, and lighter guard hairs along the edges of a dark mane and tail.
The Turkmenian kulan, also called Transcaspian wild ass, Turkmenistani onager or simply the kulan, is a subspecies of onager native to Central Asia. It was declared Endangered in 2016.