Kunga (equid)

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Kunga
Standard of Ur - War - equid.jpg
Scientific classification OOjs UI icon edit-ltr.svg
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Perissodactyla
Family: Equidae
Tribe: Equini
Genus: Equus
Species:

The kunga was a hybrid equid that was used as a draft animal in ancient Syria and Mesopotamia, where it also served as an economic and political status symbol. Cuneiform writings from as early as the mid-third millennium BCE describe the animal as a hybrid but do not provide the precise taxonomical nature of the breeding that produced it. Modern paleogenomics has revealed it to have been the offspring of a female domesticated donkey and a wild male Syrian wild ass (a subspecies of onager). They fell out of favor after the introduction of domesticated horses and mules into the region at the end of the 3rd millennium BCE.

Contents

Elite equids

Third-millennium BCE cuneiform from the kingdom of Ebla and the Mesopotamian region of Diyala name several types of equids (ANŠE, 𒀲), including one specified as the kúnga (ANŠE BAR.AN, 𒀲𒁇𒀭), which appear between about 2600 and 2000 BCE. [1] [2] These expensive animals, highly valued by the elite, [3] were purpose-bred at Nagar, the rulers of which used them themselves and monopolized their production for distribution in the region. Records from Ebla report repeated expensive purchases of kunga equids from Nagar, and it was apparently in relation to this trade that the 'high superintendents of charioteers' and those responsible for maintaining the Ebla kunga herd traveled to Nagar. [4] [5] The Ebla king gave them as gifts to other rulers. [2] It has been suggested that the kunga trade was central to the economies of the region's kingdoms, and that the ostentatious display of such expensive animals in official art directly associated them with kingship and power. [4] A pair of seals from the period, including one from Nagar, depict equids with gods in the divine realm. [4]

Hybrid nature

Contemporary descriptions of the production of the kunga seem to indicate that they were hybrids, [3] [6] and there are indications that, like most hybrid equids, they were sterile. [2] For example, foals are described in nursery herds with adult donkeys or onagers and donkey foals, never with kunga parents. [2] Production would thus have been an intensive process: they would not have established a domesticated line, but rather each individual kunga had to be produced de novo by breeding two parental species anew, [1] without the opportunity for improvement through selective breeding. Likewise, the necessity of repeated purchase of new animals from their limited production centers to maintain a stable of kunga suggests they could not be bred. [2]

Depictions

Detail from 'War' panel of the Standard of Ur mozaic, ca. 2600 BCE, showing a four-wheeled battle-wagon pulled by equids harnessed with a rein ring and a ring through the upper lip. Etendard d'Ur - char.jpg
Detail from 'War' panel of the Standard of Ur mozaic, ca. 2600 BCE, showing a four-wheeled battle-wagon pulled by equids harnessed with a rein ring and a ring through the upper lip.

Kunga were used as draft animals, with smaller males and females used for pulling plows, while 'superior' males are described in more ceremonial and martial roles, pulling the four-wheeled battle wagons and chariots of kings and gods. [2] Equids appear in this role in official imagery such as the ca. 2600 BCE Standard of Ur mozaic [2] and numerous surviving seals, [4] while a rein ring similar to those depected in the mozaic has been found at Ur, decorated with an equid. [1] These depictions are likely kunga rather than donkeys, which appear only in lesser roles in descriptions. [2] Illustrations appear to show the draft team of equids being controlled by strings passed through rings placed in the equids' upper lips. [7] Their appearance in formal administrative cuneiform and official art seems to parallel the contemporary development of kingships in the region, suggesting a propagandistic association of the kunga with royalty. [2]

Archaeology and paleogenomics

They are known to have been used for funerary purposes, as demonstrated by high-status-funeral disbursement records for harnesses, [3] and they have been identified with more than 40 equids that were sacrificed and ceremonially buried in elite graves at Umm el-Marra, Syria, in separate chambers from the burials of adult humans but many accompanied by human infants with signs of having been sacrificed. These buried kunga may have been intended either as offerings to deities, or as companions of the buried human elites, [8] while such burials may also have served a legitimizing role for the royal lines and elite, with sacrificed 'royal' equids serving as analogs of human royals. [2] Like the 'superior' kunga of cuneiform, these equids were all male, [2] ranging in size from 1.19 m to 1.36 m. [7] There are inherent challenges in identifying the species of equid skeletons, [1] [6] but the Umm el-Marra equids shared signs of domestication such as bit wear and evidence of foddering rather than grazing. [1] [3] They had a prominent overbite, while their bones had a combination of onager and donkey characteristics, being sized more like the former, but with the greater robustness of the latter, as might be expected in a hybrid between the two equid species. [1] [3] [9] Such a hybrid would have been stronger and faster than the donkey, while less intractable to taming than the Syrian wild ass. [3] [9] Their hypothesized taxonomic identity was proven by a genomic analysis reported in 2022 that compared genomes from several of these skeletons with those of extant and extinct equids, and concluded that all of the Umm el-Marra skeletons were F1 hybrid progeny of captured male Syrian wild asses with female domesticated donkeys (jenny). [1] These results make the kunga the earliest known human-engineered hybrid animal, predating the earliest mule by about 1500 years. [9] The preference for a jenny over a jack (male) as the donkey parent represents a conscious choice to have the more tractable domestic species as the maternal parent for simpler husbandry. [6] That all tested individuals were F1 hybrids reinforces the likelihood that kunga were sterile. [1]

Fate

Though the kunga held its elite status for half a millennium, it would be supplanted by both domestic horses and their donkey hybrid, the mule, introduced to the region at the end of the third millennium BCE and after that time seen filling the roles previously occupied by the kunga, [9] which rapidly disappear from the historical record. A similar hybrid was reportedly produced at the London Zoo in 1883, [10] but the subsequent extinction of the Syrian wild ass makes it impossible now to reproduce the kunga's precise taxonomic cross. [9]

Related Research Articles

The 3rd millennium BC spanned the years 3000 to 2001 BC. This period of time corresponds to the Early to Middle Bronze Age, characterized by the early empires in the Ancient Near East. In Ancient Egypt, the Early Dynastic Period is followed by the Old Kingdom. In Mesopotamia, the Early Dynastic Period is followed by the Akkadian Empire. In what is now Northwest India and Pakistan, the Indus Valley civilization developed a state society.

<span class="mw-page-title-main">Hybrid (biology)</span> Offspring of cross-species reproduction

In biology, a hybrid is the offspring resulting from combining the qualities of two organisms of different varieties, species or genera through sexual reproduction. Generally, it means that each cell has genetic material from two different organisms, whereas an individual where some cells are derived from a different organism is called a chimera. Hybrids are not always intermediates between their parents, but can show hybrid vigor, sometimes growing larger or taller than either parent. The concept of a hybrid is interpreted differently in animal and plant breeding, where there is interest in the individual parentage. In genetics, attention is focused on the numbers of chromosomes. In taxonomy, a key question is how closely related the parent species are.

<span class="mw-page-title-main">Domestication of the horse</span>

How and when horses became domesticated has been disputed. Although horses appeared in Paleolithic cave art as early as 30,000 BC, these were wild horses and were probably hunted for meat. The clearest evidence of early use of the horse as a means of transport is from chariot burials dated c. 2000 BC. However, an increasing amount of evidence began to support the hypothesis that horses were domesticated in the Eurasian Steppes in approximately 3500 BC. Discoveries in the context of the Botai culture had suggested that Botai settlements in the Akmola Province of Kazakhstan are the location of the earliest domestication of the horse. Warmouth et al. (2012) pointed to horses having been domesticated around 3000 BC in what is now Ukraine and Western Kazakhstan.

<span class="mw-page-title-main">Donkey</span> Domesticated animal used for transportation

The donkey is a domesticated equine. It derives from the African wild ass, Equus africanus, and may be classified either as a subspecies thereof, Equus africanus asinus, or as a separate species, Equus asinus. It was domesticated in Africa some 5000–7000 years ago, and has been used mainly as a working animal since that time.

<span class="mw-page-title-main">Onager</span> Species of mammal

The onager, also known as hemione or Asiatic wild ass, is a species of the family Equidae native to Asia. A member of the subgenus Asinus, the onager was described and given its binomial name by German zoologist Peter Simon Pallas in 1775. Six subspecies have been recognized, two of which are extinct.

<span class="mw-page-title-main">Ebla</span> Ancient Syrian city

Ebla was one of the earliest kingdoms in Syria. Its remains constitute a tell located about 55 km (34 mi) southwest of Aleppo near the village of Mardikh. Ebla was an important center throughout the 3rd millennium BC and in the first half of the 2nd millennium BC. Its discovery proved the Levant was a center of ancient, centralized civilization equal to Egypt and Mesopotamia and ruled out the view that the latter two were the only important centers in the Near East during the Early Bronze Age. The first Eblaite kingdom has been described as the first recorded world power.

Umm el-Marra,, east of modern Aleppo in the Jabbul Plain of northern Syria, was one of the ancient Near East's oldest cities, located on a crossroads of two trade routes northwest of Ebla, in a landscape that was much more fertile than it is today. Possibly this is the city of Tuba mentioned in Egyptian inscriptions listing cities that were defeated or destroyed in the Pharaoh Thutmose III's north Syrian campaign. The city of Tuba is also mentioned in epigraphic remains from Ebla, Mari, and Alalakh.

<span class="mw-page-title-main">Zebroid</span> Offspring of a zebra and any other equine

A zebroid is the offspring of any cross between a zebra and any other equine to create a hybrid. In most cases, the sire is a zebra stallion. The offspring of a donkey sire and zebra dam, called a donkra, and the offspring of a horse sire and a zebra dam, called a hebra, do exist, but are rare and are usually sterile. Zebroids have been bred since the 19th century. Charles Darwin noted several zebra hybrids in his works.

<i>Equus</i> (genus) Genus of mammals which includes horses, donkeys, and zebras

Equus is a genus of mammals in the family Equidae, which includes horses, asses, and zebras. Within the Equidae, Equus is the only recognized extant genus, comprising seven living species. Like Equidae more broadly, Equus has numerous extinct species known only from fossils. The genus most likely originated in North America and spread quickly to the Old World. Equines are odd-toed ungulates with slender legs, long heads, relatively long necks, manes, and long tails. All species are herbivorous, and mostly grazers, with simpler digestive systems than ruminants but able to subsist on lower-quality vegetation.

<span class="mw-page-title-main">Kiang</span> Tibetan wild ass

The kiang is the largest of the Asinus subgenus. It is native to the Tibetan Plateau in Ladakh, northern Pakistan, Tajikistan, China and northern Nepal. It inhabits montane grasslands and shrublands. Other common names for this species include Tibetan wild ass, khyang and gorkhar.

<span class="mw-page-title-main">Domestication of vertebrates</span>

The domestication of vertebrates is the mutual relationship between vertebrate animals including birds and mammals, and the humans who have influence on their care and reproduction.

<span class="mw-page-title-main">Tell Brak</span> Archaeological site in Syria

Tell Brak was an ancient city in Syria; its remains constitute a tell located in the Upper Khabur region, near the modern village of Tell Brak, 50 kilometers north-east of Al-Hasaka city, Al-Hasakah Governorate. The city's original name is unknown. During the second half of the third millennium BC, the city was known as Nagar and later on, Nawar.

<span class="mw-page-title-main">African wild ass</span> Species of wild ass

The African wild ass or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey, which is sometimes placed within the same species. They live in the deserts and other arid areas of the Horn of Africa, in Eritrea, Ethiopia and Somalia. It formerly had a wider range north and west into Sudan, Egypt, and Libya. It is Critically Endangered, with about 570 existing in the wild.

<i>Asinus</i> Subgenus of mammals

Asinus is a subgenus of Equus that encompasses several subspecies of the Equidae commonly known as wild asses, characterized by long ears, a lean, straight-backed build, lack of a true withers, a coarse mane and tail, and a reputation for considerable toughness and endurance.

<span class="mw-page-title-main">Early Dynastic Period (Mesopotamia)</span> Archaeological culture of Mesopotamia

The Early Dynastic period is an archaeological culture in Mesopotamia that is generally dated to c. 2900 – c. 2350 BC and was preceded by the Uruk and Jemdet Nasr periods. It saw the development of writing and the formation of the first cities and states. The ED itself was characterized by the existence of multiple city-states: small states with a relatively simple structure that developed and solidified over time. This development ultimately led to the unification of much of Mesopotamia under the rule of Sargon, the first monarch of the Akkadian Empire. Despite this political fragmentation, the ED city-states shared a relatively homogeneous material culture. Sumerian cities such as Uruk, Ur, Lagash, Umma, and Nippur located in Lower Mesopotamia were very powerful and influential. To the north and west stretched states centered on cities such as Kish, Mari, Nagar, and Ebla.

<span class="mw-page-title-main">Syrian wild ass</span> Extinct subspecies of onager

The Syrian wild ass, less commonly known as a hemippe, an achdari, or a Mesopotamian or Syrian onager, is an extinct subspecies of onager native to the Arabian peninsula and surrounding areas. It ranged across present-day Iraq, Palestine, Israel, Iran, Jordan, Saudi Arabia, Syria, and Turkey.

<span class="mw-page-title-main">Turkmenian kulan</span> Subspecies of onager

The Turkmenian kulan, also called Transcaspian wild ass, Turkmenistani onager or simply the kulan, is a subspecies of onager native to Central Asia. It was declared Endangered in 2016.

<span class="mw-page-title-main">The Dog and the Wolf</span> Aesops fable

The Dog and the Wolf is one of Aesop's Fables, numbered 346 in the Perry Index. It has been popular since antiquity as an object lesson of how freedom should not be exchanged for comfort or financial gain. An alternative fable with the same moral concerning different animals is less well known.

<span class="mw-page-title-main">Persian onager</span> Subspecies of onager

The Persian onager, also called the Persian wild ass or Persian zebra, is a subspecies of onager native to Iran (Persia). It is listed as Endangered, with no more than 600 individuals left in the wild and only 30 individuals living within North American institutions.

Equid hybrids, also called hybrid equines, are created from the crossing of members from the horse family such as a horse, donkey and zebra.

References

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