Biokoviella | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Subphylum: | Myriapoda |
Class: | Diplopoda |
Order: | Chordeumatida |
Family: | Anthogonidae |
Subfamily: | Biokoviellinae |
Genus: | Biokoviella Mršić, 1992 |
Biokoviella is a genus of millipedes belonging to family Anthogonidae in order Chordeumatida. [1] Adult millipedes have 30 segments (counting the collum as the first segment and the telson as the last). [2]
The genus contains two recognised species: [1]
The genus was formerly assigned to the monotypic family Biokoviellidae, [2] but is now treated as a monotypic subfamily in family Anthogonidae. [1]
Glomerida is an order of pill-millipedes found primarily in the Northern Hemisphere. Also known as northern pill millipedes, they superficially resemble pill-bugs or woodlice, and can enroll into a protective ball. They have twelve body segments, 17 to 19 pairs of legs, and males have enlarged rear legs involved in mating. The order includes about 30 genera and at least 280 species, including Glomeris marginata, the common European pill-millipede. The order contains members in Europe, South-east Asia and the Americas from California to Guatemala. Although historically considered closely related with the similar sphaerotheriidans that also enroll, some DNA evidence suggest they may be more closely related to glomeridesmidans, a poorly known order that does not enroll.
Polydesmida is the largest order of millipedes, with more than 5,000 species, including all the millipedes reported to produce hydrogen cyanide (HCN). This order is also the most diverse of the millipede orders in terms of morphology. Millipedes in this order are found in all regions of the world other than Antarctica.
Xystodesmidae is a family of millipedes in the suborder Leptodesmidea within the order Polydesmida. The family Xystodesmidae was created by the American biologist Orator F. Cook in 1895 and named after the genus Xystodesmus. This family includes more than 390 known species distributed among 62 genera. Many species, however, remain undescribed: for example, it is estimated that the genus Nannaria contains over 200 species, but only 25 were described as of 2006. By 2022, 78 species in Nannaria have been described.
Zephroniidae is a family of giant pill millipedes in the taxonomic order Sphaerotheriida. They occur in southeast Asia from the Himalayas and China south and east to Sulawesi and to Australia, and also inhabit some Philippine islands.
Polyxenida is an order of millipedes readily distinguished by a unique body plan consisting of a soft, non-calcified body ornamented with tufts of bristles. These features have inspired the common names bristly millipedes or pincushion millipedes. This order includes about 148 species in four families worldwide, which represent the only living members of the subclass Penicillata.
Polyzoniida is an order of millipedes in the subterclass Colobognatha. This order contains three families and more than 70 described species. The species in this order are also known as camphor millipedes, because ozopore secretions in this order frequently have a strong camphor-like smell. Poison frogs in South America and Madagascar evidently obtain some of their poison from these millipedes.
Glomeridesmida is an order of millipedes in the infraclass Pentazonia containing two families and at least 35 described species. Glomeridesmida is the only living order of the superorder Limacomorpha. Glomeridesmidans are also known as slug millipedes.
Siphonophorida is an order of millipedes containing two families and over 100 species.
Chordeumatida is a large order of millipedes containing more than 1,400 species. Also known as sausage millipedes, they are found nearly worldwide. Chordeumatida is the largest order in the superorder Nematophora, a group also known as spinning millipedes because their telsons feature spinnerets used to build nests of silk. These millipedes produce this silk to create chambers in which to molt or to lay their eggs.
Urochordeuma is a genus of millipedes in the order Chordeumatida and the only genus in the family Urochordeumatidae. Adult millipedes in this family have 30 segments. The two species are known from the U.S. state of Washington, where U. bumpusi is known from Pierce County, and U. porona from King County.
Siphoniulus is a poorly known genus of millipede containing only two living species: S. alba from Indonesia, and S. neotropicus from Mexico and Guatemala. An additional two fossil species are known from Cretaceous amber. Siphoniulus species are the only members of the family Siphoniulidae and order Siphoniulida, making Siphoniulida the smallest millipede order. Few specimens are known, and their classification is contentious, although most recent studies place them as basal members of the Helminthomorpha.
Haplogona is a genus of chordeumatidan millipedes and the only genus in the family Verhoeffiidae. Adult millipedes in this family have 30 segments. These species occur in the southern Alps of Europe, from Genoa, Italy to the Istrian peninsula.
Rhiscosomides is a genus of millipedes in the order Chordeumatida with seven described species, and is the only genus in the family Rhiscosomididae. Adult millipedes in this family have 30 segments.
Kirkayakidae is a family of millipedes belonging to the order Chordeumatida. This family was formerly known as Altajellidae. Adult millipedes in this family have 28 segments rather than the 30 segments usually found in chordeumatidans.
Hoffmaneumatidae is a family of millipedes belonging to the order Chordeumatida. This family includes two genera, one (Hoffmaneuma) found in the Russian Far East and the other (Japanoparvus) found in Japan. Millipedes in this family range from 4 mm to 6 mm in length. Adult millipedes in this family have only 28 segments, not the 30 segments usually found in this order. Adult males in this family feature a reduced leg pair 10 in addition to the two pairs normally modified into gonopods in this order. In the adult male of the species Hoffmaneuma exiguum, for example, the gonopod complex derives from all three leg pairs rather than from just the usual two.
Entomobielziidae is a family of millipedes belonging to the order Chordeumatida. This family includes 18 species, including 13 in the genus Tianella. Millipedes in this family are found in Romania, central Asia, and the Himalayas.
Kashmireumatidae is a small family of millipedes belonging to the order Chordeumatida. These millipedes range from 5mm to 14mm in length and are found in East Asia. Adult millipedes in this family have only 26 or 28 segments rather than the 30 segments usually found in this order. In the species Vieteuma topali, adults have 26 segments, whereas in all other species in this family, adults have 28 segments.
Chamaesoma is a monotypic genus of millipede in the family Chamaesomatidae, and Chamaesoma broelemanni is the only species in this genus. This millipede is notable as one of only a few species in the order Chordeumatida with only 26 segments in adults, four fewer segments than typically found in adults this order. This species lives on the surface of the soil and is distributed across most of France and Luxembourg. The genus Chamaesoma and its only species were first described by the German zoologist Karl W. Verhoeff in 1913 based on type material found by the French zoologists Henri Brölemann and Henri Ribaut under wet leaf litter near St. Béat in the Pyrenees in France.
Metamastigophorophyllon is a genus of millipedes in the family Anthroleucosomatidae. This genus includes millipedes found in Azerbaijan, Iran, Russia, and Georgia. All five species in this genus have 31 segments in adults. This genus is notable for including the only species in the order Chordeumatida with 31 segments rather than the 30 segments usually observed in adults in this order.
Agenodesmus reticulatus is a species of millipede in the family Fuhrmannodesmidae, which some authorities consider a junior synonym of Trichopolydesmidae. This millipede is among the very few species in the order Polydesmida to feature adults with only 18 segments rather than the 20 segments usually found in this order. This species is notable as the first polydesmidan millipede discovered with only 18 segments in adults, the smallest number recorded in the order Polydesmida. Before the discovery of A. reticulatus, polydesmidans were known to have only 19 or 20 segments in adults.