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The siphuncle is a strand of tissue passing longitudinally through the shell of a cephalopod mollusk. Only cephalopods with chambered shells have siphuncles, such as the extinct ammonites and belemnites, and the living nautiluses, cuttlefish, and Spirula. In the case of the cuttlefish, the siphuncle is indistinct and connects all the small chambers of that animal's highly modified shell; in the other cephalopods it is thread-like and passes through small openings in the septa (walls) dividing the camerae (chambers). Some older studies have used the term siphon for the siphuncle, though this naming convention is uncommon in modern studies to prevent confusion with a mollusc organ of the same name.
Nautiloids are a group of marine cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living Nautilus and Allonautilus. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms (orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day.
Cameroceras is an extinct genus of endocerid cephalopod which lived in equatorial oceans during the entire Ordovician period. Like other endocerids, it was an orthocone, meaning that its shell was fairly straight and pointed. It was particularly abundant and widespread in the Late Ordovician, inhabiting the shallow tropical seas in and around Laurentia, Baltica and Siberia.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
Mcqueenoceras is an extinct genus of early endocerid, a nautiloid from the Floian epoch of the late early Ordovician period. It was similar in overall form to Clitendoceras, from which it may have been derived. Mcqueenoceras, like Clitendoceras, has ventral siphuncle but the endocones are thicker on the ventral side and thinner on the dorsal. Also the sutures in Mcqueenoceras retreat rearward, forming lobes as they cross the venter. The type species is Mcqueenoceras jeffersonense, named by E.O. Ulrich and A.F. Foerste in 1935, and it is known from Missouri and New York. In 1956, Rousseau H. Flower named two other species, M. cariniferum and M. ventrale, both known from Maryland.
Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.
Cyptendoceras is an extinct nautiloid cephalopod included in the family Ellesmeroceratidae that lived in what would be North and South America during the latter part of the Early Ordovician from about 475 – 472 mya, existing for approximately 3 million years.
Cyrtobaltoceras is an extinct cephalopod genus known from the upper Lower Ordovician Fort Cassin Formation at Valcour, N.Y. that is included in the Nautiloid family Baltoceratidae
Rioceras is an extinct genus of orthocerid cephalopods of the family Baltoceratidae that lived in what would be North America during the Early Ordovician from 480—472 mya, existing for approximately 8 million years.
Rhabdiferoceras is an extinct genus of orthocerids belonging to the Baltoceratidae that lived in what would be North America during the Cassinian Stage at the end of the Early Ordovician, existing for approximately two million years from about 474 -472 mya.
Microbaltoceras is an extinct genus of the cephalopod family Baltoceratidae that lived in what would be North America during the Early Ordovician. It was named by Rousseau Flower (1964) and assigned by him to the Baltoceratidae. The type species is Microbaltoceras minore which was found in the Threadgill Member of the Tanyard Formation in Gillespie County, Texas, USA.
Metabaltoceras, first described by Rousseau Flower in 1964, is a fossil cephalopod genus in the family Baltoceratidae, with a small, slender, fairly fusiform shell, and a large empty siphuncle in contact with the ventral surface. The siphuncle is achoanitic, having virtually no septal necks, segments formed all but entirely by connecting rings. Shells are generally straight, beginning with a subcircular cross section in the adapical portion but becoming faintly depressed adorally. Sutures are straight and transverse except ventrally, where they produce deep, prominent lobes.
Orthoceratoidea is a major subclass of nautiloid cephalopods. Members of this subclass usually have orthoconic (straight) to slightly cyrtoconic (curved) shells, and central to subcentral siphuncles which may bear internal deposits. Orthoceratoids are also characterized by dorsomyarian muscle scars, extensive cameral deposits, and calciosiphonate connecting rings with a porous and calcitic inner layer.
The Graciloceratidae is a family of nautiloid cephalopods from the Middle and Upper Ordovician belonging to the Oncocerida, characterized by exogastric cyrtocones that expand slightly or moderately and have thin walled, orthochoanitic marginal or subventral, tubular siphuncles.
Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.
Tajaroceras is an extinct slender cephalopod from the uppermost Lower Ordovician of western North America, belonging to the Orthocerid family Troedssonellidae.
Veneficoceras is a genus of the rod-bearing Baltoceratidae, an extinct cephalopod family with characteristics of the orthoceratoid Dissidocerida, found in Cassinianage, Lower Ordovician, limestone in western Utah.
The Sactorthoceratidae comprise Orthocerataceaen genera with a subcentral suborthochoanitic siphuncle composed of slightly expanded segments and free of organic deposits. The camerae (chambers) of the phragmocone likewise have organic deposits that are typically retarded or sparse.
Murrayoceras is a nautilid cephalopod included in the orthocerid family Baltoceratidae, widespread in the Middle Ordovician of North America, characterized by a depressed orthoconic shell with a subtriangular cross section and flattened venter and a proportionally large ventral siphuncle, 0.15 to 0.3 the dorso-ventral shell diameter. Septa are close spaced with sutures forming broad lobes on the upper flanks and ventral surface.
Apocrinoceratidae constitutes a family of Middle Ordovician nautiloid cephalopods characterized by straight or slightly curved, transversely ribbed shells having siphuncles composed of expanded segments, short recurved septal necks, and thick connecting rings. Derivation is from the Protocycloceratidae, a family of ellesmerocerids, which differ in having straight or concave siphuncle segments, but are otherwise similar in form.
References
- 1 2 3 Rousseau H Flower, (1964). The nautiloid order Ellesmeroceratida (Cephalopoda). New Mexico Bureau of Mines and Mineral Resources Memoir 12.
- ↑ R. C. Frey. 1995. Middle and Upper Ordovician nautiloid cephalopods of the Cincinnati Arch region of Kentucky, Indiana, and Ohio. U.S. Geological Survey Professional Paper 1066P:1-126
- ↑ B. Kröger, et al . 2007. Early orthoceratoid cephalopods from the Argentine Precordillera (Lower-Middle Ordovician). Journal of Paleontology 81(6):1266-1283