Cixiinae | |
---|---|
Cixius nervosus | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Arthropoda |
Class: | Insecta |
Order: | Hemiptera |
Suborder: | Auchenorrhyncha |
Infraorder: | Fulgoromorpha |
Family: | Cixiidae |
Subfamily: | Cixiinae Spinola, 1839 |
Tribes | |
See text |
Cixiinae is a planthopper subfamily in the family Cixiidae. It is one of three such subfamilies, the other two being the Bothriocerinae and the Borystheninae. [1] While a few species had been tested in a larger study of the Fulgoroidea, [2] neither the Cixiinae nor its tribes were analysed cladistically until 2002. [1] [3] Resolution of tribal relationships is incomplete and additional testing of the tribes with samples larger than one per tribe is needed. [4]
In 1938 Metcalf divided the Cixiinae into tribes, including the Cixiini and Oecleini of Muir; [5] however, those divisions were seldom referred to and remained unanalysed for decades. [2] Subsequently, the Stenophlepsiini were returned to the Cixiinae, and the Pentastirini were subdivided into the Pentastirina and Oliarina subtribes. [2] [6] However, Oliarina was later synonymized under Pentastirina. [7] In 2002, Emeljanov created seven new tribes [3] [4] and in 2004 subtribe Mnemosynina of tribe Pentastirini was raised to tribal level as the Mnemosynini. [8]
Currently there are sixteen recognized tribes, although some tribes lack adequate description: [4]
The Auchenorrhyncha suborder of the Hemiptera contains most of the familiar members of what was called the "Homoptera" – groups such as cicadas, leafhoppers, treehoppers, planthoppers, and spittlebugs. The aphids and scale insects are the other well-known "Homoptera", and they are in the suborder Sternorrhyncha.
A planthopper is any insect in the infraorder Fulgoromorpha, in the suborder Auchenorrhyncha, a group exceeding 12,500 described species worldwide. The name comes from their remarkable resemblance to leaves and other plants of their environment and that they often "hop" for quick transportation in a similar way to that of grasshoppers. However, planthoppers generally walk very slowly. Distributed worldwide, all members of this group are plant-feeders, though few are considered pests. Fulgoromorphs are most reliably distinguished from the other Auchenorrhyncha by two features; the bifurcate (Y-shaped) anal vein in the forewing, and the thickened, three-segmented antennae, with a generally round or egg-shaped second segment (pedicel) that bears a fine filamentous arista.
The Cixiidae are a family of fulgoroid insects, one of many families commonly known as planthoppers, distributed worldwide and comprising more than 2,000 species from over 150 genera.
The family Derbidae is a large and diverse group of planthoppers, widely distributed in the tropics and also found in subtropical and temperate regions. It includes more than 150 genera and more than 1500 species. The adults suck the sap of plants while the nymphs live mainly in decaying organic matter, for example under the bark of dead tree trunks, feeding on fungi. In some groups of species the general structure of the insects is largely uniform, whereas in others like in the subfamily Otiocerinae various forms can be found. For example, the shape of the head, the antennae or the wings can differ considerably among genera and species.
Dictyopharidae is a family of planthoppers, related to the Fulgoridae. The family comprises nearly 760 species in more than 150 genera which are grouped into two subfamilies, Dictyopharinae and Orgeriinae.
Flatidae are a family of fulgoroid planthoppers. They are cosmopolitan in distribution and are distinguished from others in the superfamily by a combination of characters. Like all other planthoppers, they suck phloem sap of plants. Some species are known to communicate with vibrations through the plant stems. Communication may be with mates, or with ants that tend the nymphs, protecting them and gathering honeydew secretions. Adults of some species have brightly coloured forewings which are tougher and known as tegmina unlike the membranous hindwings which are used for flight. Although a few can be identified by their coloration, most species requires dissection and examination under a microscope with access to literature on already described species.
Issidae is a family of planthoppers described by Spinola in 1839, belonging to the order Hemiptera, suborder Auchenorrhyncha superfamily Fulgoroidea.
Nogodinidae is a family of planthoppers. They have membranous wings with delicate venation and can be confused with members of other Fulgoroid families such as the Issidae and Tropiduchidae. Some authors treat it as a subfamily of the Issidae.
Lophopidae is a family of fulgoroid plant-hoppers with most species found in tropical South America and Asia.
Cixiini is a planthopper tribe in the family Cixiidae. This tribe is non-monophyletic.
Melanoliarus is a genus of cixiid planthoppers in the family Cixiidae. There are at about 50 described species in Melanoliarus, which are common and widespread in the Nearctic and Neotropics.
Achilidae is a family of planthoppers, sometimes called "achilids" in the order Hemiptera. There are at least 520 described species in Achilidae.
Otiocerinae is a subfamily of planthoppers in the family Derbidae with more than 100 genera and more than 1000 species. The species are widely distributed, especially in the Americas, Sub-Saharan Africa, as well as parts of Asia, Australia and the Pacific. Like in other Derbidae, the adults suck the sap of plants while the nymphs live mainly in decaying organic matter, for example under the bark of dead tree trunks, feeding on fungi.
Asiracinae is a subfamily of delphacid planthoppers in the family Delphacidae. There are at least 30 genera and 180 described species in Asiracinae, which probably has a world-wide distribution.
Tangiini is a tribe of tropiduchid planthoppers in the subfamily Tropiduchinae. There are at least 30 described species in Tangiini, from the Americas with several genera apparently restricted to the Caribbean.
Tropiduchinae is a subfamily of tropiduchid planthoppers in the family Tropiduchidae.
Derbinae is a subfamily of derbid planthoppers in the family Derbidae.
Dictyopharinae is a subfamily of dictyopharid planthoppers in the family Dictyopharidae. There more than 100 genera and 500 described species in Dictyopharinae.
The Achilixiidae are a family of Fulgoromorpha (planthoppers); species may be found in the neotropical and Asian regions. They are closely related to Achilidae and are sometimes included under Achilidae as a subfamily. Like Achilidae, species generally feed on several species of plant though the nymph stage has been found to feed on fungus. Like other planthoppers, the immature stage is covered in a wax which may help protect it from predators. Achilixiidae are small or medium sized for planthoppers and are greatly compressed, not depressed like the Achilidae.
Perforissidae is an extinct family of planthoppers. They are considered to belong to the group of "Cixiidae-like" planthoppers. Species are known from the Early to Late Cretaceous of Eurasia, North America and South America. The family was named by Shcherbakov in 2007