Cloacaspis

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Cloacaspis
Temporal range: Early Arenig [1]
Cloacaspis.jpg
C. senilis, and C. ekphymosa
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Cloacaspis

Fortey 1974 [2]

Cloacaspis is an extinct genus of Olenid Ptychopariid trilobite. It lived during the early part of the Arenig stage of the Ordovician Period, [1] a faunal stage which lasted from approximately 478 to 471 million years ago. Richard Fortey has proposed that these particular trilobites lived in anoxic regions of the ocean floor, and cultivated symbiotic, sulfur-metabolizing bacteria. [3]

A genus is a taxonomic rank used in the biological classification of living and fossil organisms, as well as viruses, in biology. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus.

The Olenidae were a family of trilobites. They are thought to have evolved a symbiotic relationship with sulfur-eating bacteria from which they derived food.

Ptychopariida order of arthropods (fossil)

Ptychopariida is a large, heterogeneous order of trilobite containing some of the most primitive species known. The earliest species occurred in the second half of the Lower Cambrian, and the last species did not survive the Ordovician–Silurian extinction event.

Etymology

The generic name is a compound word from Latin, "cloaca," meaning sewer, and from Greek, "aspis," meaning "shield." Thus, Cloacaspis translate as "sewer shield." In his book, Trilobite! Eyewitness to Evolution, Fortey, who first discovered the genus in Ordovician strata in Spitsbergen, explained that the rocks they were found in had a rank, sulfurous odor, reminding him of raw sewage. [4]

Related Research Articles

Trilobite class of arthropods (fossil)

Trilobites are a group of extinct marine arachnomorph arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period, and they flourished throughout the lower Paleozoic era before beginning a drawn-out decline to extinction when, during the Devonian, all trilobite orders except the Proetids died out. Trilobites disappeared in the mass extinction at the end of the Permian about 252 million years ago. The trilobites were among the most successful of all early animals, existing in oceans for almost 300 million years.

Richard Fortey British paleontologist

Richard Alan Fortey FRS FRSL is a British palaeontologist, natural historian, writer and television presenter, who served as President of the Geological Society of London for its bicentennial year of 2007; he is married and has four children.

Svalbardites is an extinct genus of trilobites. It lived during the Arenig stage of the Ordovician Period, approximately 478 to 471 million years ago.

<i>Balnibarbi</i> (trilobite) genus of arthropods (fossil)

Balnibarbi is an extinct genus of trilobites in the family Olenidae. They are known from fossils excavated in Norway. They lived during the early part of the Arenig stage of the Ordovician Period, a faunal stage that occurred about 479 to 472 million years ago.

<i>Globampyx</i> genus of arthropods (fossil)

Globampyx is an extinct genus raphiophorid trilobites. It lived during the later part of the Arenig stage of the Ordovician Period, approximately 478 to 471 million years ago. Species of the genus are known from Canada, Norway (Svalbard) and Sweden.

<i>Pytine</i> genus of trilobites

Pytine is an extinct genus of asaphid trilobites. Species lived during the later part of the Arenig stage of the Ordovician Period, approximately 478 to 471 million years ago. Various species are found in the Svalbard, Valhallfonna Formation, Olenidsletta, Member, of Spitzbergen, Norway, the Megistaspis (Paramegistaspis) planilimbata Zone of the 'Shumardia Shale' of Sweden, Jujuy Province, Argentina, early Arenig-aged strata of Jiangxi province, China, and Darriwilian-aged strata in Western Hunan province, China. The type species, P. graia, has seven thorax segments, and lacks the rapier-like glabellar spine, that occurs in many other raphiophorids. The Chinese species, by contrast, have only six thoracic segments. So far, only the type species, and one of the Chinese species, P. laevigata, are known from complete specimens.

Thysanopeltis is a genus of trilobite that lived from the Early to the Middle Devonian. Its remains have been found in Africa and Europe.

<i>Crotalocephalus</i> genus of trilobites

Crotalocephalus is a genus of trilobites that lived from the Late Silurian to the Early Devonian. Its remains have been found in Asia, Australia, Africa, and Europe.

<i>Cheirurus</i> genus of trilobites

Cheirurus is a genus of phacopid trilobites that lived from the Late Cambrian to the Middle Devonian. Its remains have been found in Africa, Asia, Australia, Europe, and North America. Cheirurus is the type genus of Cheiruridae.

<i>Cyphaspis</i> genus of trilobites

Cyphaspis is a genus of small trilobite that lived from the Late Ordovician to the Late Devonian. Fossils have been found in marine strata in what is now Europe, Africa and North America. Various species had a compact body, and a large, bulbous glabellum. Many species had long spines arranged similarly to closely related genera, such as Otarian, Otarionella, Chamaeleoaspis, and Namuropyge.

<i>Norasaphus monroeae</i> species of trilobite

Norasaphus monroeae is a species of asaphid trilobites named after Marilyn Monroe for its hourglass-like shaped glabellum. Its fossils are found in Arenig-aged marine strata from the Nora Formation, in the Georgina Basin, situated between the Northern Territory and Queensland, Australia.

Annamitella Mansuy 1916 is a genus of trilobite, extinct marine arthropods. Annamitella lived from the Arenig to the Llandeilo age of the Ordovician Period from 478.6 to 460.9 million years ago.

<i>Trinodus</i> genus of trilobites (fossil)

Trinodus is a very small to small blind trilobite, a well known group of extinct marine arthropods, which lived during the Ordovician, in what are now the Yukon Territories, Virginia, Italy, Czech Republic, Poland, Denmark, Sweden, Svalbard, Ireland, Scotland, Wales, Iran, Kazakhstan and China. It is one of the last of the Agnostida order to survive.

<i>Ogygiocarella</i> genus of trilobites

Ogygiocarella is a genus of asaphid trilobites. It occurred during the Middle Ordovician.

Prospectatrix is a genus of trilobites of average size, that lived in the Lower Ordovician and is probably ancestral to the other genera of the Cyclopygidae family. Its eyes are only moderately enlarged and it has six or seven thorax segments.

Carolinites is a genus of trilobite, assigned to the Telephinidae family, that occurs during the Lower and Middle Ordovician. Carolinites had a pantropical distribution, and there is evidence that it lived in upper parts of the water column. The free cheeks of Carolinites are largely covered by its huge eyes, except for the attachment of large genal spines that extend downward, backward and lateral and gradually curving further backward. The glabella is slightly bulbous, the occipital ring is well defined, but further transglabellar furrows are lacking. The thorax has 10 segments. The axis of the pygidium is highly vaulted, with a curved spine emerging almost perpendicular to the midline and ending parallel to it and a node on each of the other three segments. Carolinites is known from what are today Australia (Tasmania), Canada (Alberta), China, France, Spitsbergen, and the United States (Utah).

Raphiophoridae family of arthropods (fossil)

Raphiophoridae is a family of small to average-sized trilobites that first occurred at the start of the Ordovician and became extinct at the end of the Middle Silurian.

<i>Gog</i> (trilobite)

Gog is a genus of large, flattened asaphid trilobite from the Middle Arenig-aged Svalbard, Valhallfonna Formation, Olenidsletta, Member, of Spitzbergen, Norway, and the Upper Arenig-aged Dawan Formation in Hubei, China.

<i>Paraceraurus</i> genus of trilobites

Paraceraurus is a genus of trilobites that lived in the Ordovician period. Its remains have been found in China, Estonia, Sweden and North America. These trilobites have a rounded and moderately convex cephalon. Glabella is convex or flattened, with a sub-rectangular outline. Thorax shows eleven segments.

<i>Bulbaspis</i> genus of trilobites

Bulbaspis is a late Ordovician genus of asaphid trilobites of the family Raphiophoridae found primarily in upper Ordovician-aged deepwater marine strata of Kazakhstan, China, and possibly Tasmania. Species of Bulbaspis are similar to other raphiophorids such as Ampyx and Raphiophorus, save that the long spine that emanates from the glabella of the latter two genera has been modified into a knob-like or bulb-like structure in Bulbaspis that developed incrementally in the animal's growth. The function of the bulb is as yet unknown: one hypothesis suggests sexual selection may have had a role in its evolution in the genus.

References

  1. 1 2 Sepkoski, Jack (2002). "A compendium of fossil marine animal genera (Trilobita entry)". Bulletins of American Paleontology. 364: 560. Archived from the original on 2006-09-05. Retrieved 2008-01-12.
  2. Fortey, R.A., "The Ordovician Trilobites of Spitsbergen"
  3. Fortey, Richard. Trilobite! Eyewitness to Evolution Random House Inc, 2001 p 70
  4. Fortey, Richard. Trilobite! Eyewitness to Evolution Random House Inc, 2001 p 69