| Forcipomyia squamipennis | |
|---|---|
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Diptera |
| Family: | Ceratopogonidae |
| Genus: | Forcipomyia |
| Species: | F. squamipennis |
| Binomial name | |
| Forcipomyia squamipennis | |
Forcipomyia squamipennis is a biting midge species in the genus Forcipomyia . It is an important pollinator of cacao trees ( Theobroma cacao ) in Ghana, and other related midges pollinate the tree in other parts of the tropics.
Forcipomyia squamipennis is an important pollinator of the cacao tree Theobroma cacao in Ghana, [1] not because it pollinates more effectively than other insects but because it is so numerous in cacao plantations. The population is greatest in the rainy season. Adult midges spend the day in shady spots such as between the buttress roots of large trees, in crevices in logs, in hollow stumps or in piles of husk debris. They emerge at variable times of day to swarm near their hiding locations, and disperse in the late afternoons and early mornings. Most midges do not move further than about 6 m (20 ft). The females lay batches of up to about ninety eggs on damp piles of plant debris. The eggs hatch after two or three days and the larvae pass through four instar stages before pupating at about twelve days. The adults survive for about a week and there are thought to be about twelve generations of the midge per year. Both sexes feed on the pollen of the cacao flower, but four times more males visit the flowers than do females. [1]
Midges in the family Ceratopogonidae are believed to be the most important pollinators of cacao globally, particularly the larger members of the family. The tree flowers profusely but few flowers set, particularly in the dry season; when the tree is under water stress all flowers are dropped within about five days. Successful pollination requires the deposition of at least 35 suitable pollen grains on the receptive parts of the flower, and is dependent on the season. [2]
The cocoa bean or simply cocoa, also called cacao, is the dried and fully fermented seed of Theobroma cacao, from which cocoa solids and cocoa butter can be extracted. Cocoa beans native to the Ecuadorian Amazon rainforest are the basis of chocolate and Mesoamerican foods including tejate, an indigenous Mexican drink.
Theobroma cacao is a small evergreen tree in the family Malvaceae. Its seeds, cocoa beans, are used to make chocolate liquor, cocoa solids, cocoa butter and chocolate. Native to the tropics of the Americas, the largest producer of cocoa beans in 2018 was Ivory Coast, at 2.2 million tons. Its leaves are alternate, entire, unlobed, 10–50 cm (4–20 in) long and 5–10 cm (2–4 in) broad.
A pollinator is an animal that moves pollen from the male anther of a flower to the female stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains.
Pollination is the transfer of pollen from an anther of a plant to the stigma of a plant, later enabling fertilisation and the production of seeds, most often by an animal or by wind. Pollinating agents can be animals such as insects, for example beetles; birds, butterflies, and bats; water; wind; and even plants themselves. Pollinating animals travel from plant to plant carrying pollen on their bodies in a vital interaction that allows the transfer of genetic material critical to the reproductive system of most flowering plants. When self-pollination occurs within a closed flower. Pollination often occurs within a species. When pollination occurs between species, it can produce hybrid offspring in nature and in plant breeding work.
Theobroma grandiflorum, commonly known as cupuaçu, also spelled cupuassu, cupuazú, cupu assu, or copoazu, is a tropical rainforest tree related to cacao. Native and common throughout the Amazon basin, it is naturally cultivated in the jungles of north of Brazil, with the largest production in Pará, Amazonas and Amapá, Colombia, Bolivia and Peru. The pulp of the cupuaçu fruit is consumed throughout Central and South America, especially in the northern states of Brazil, and is used to make ice creams, snack bars, and other products.
A midge is any small fly, including species in several families of non-mosquito Nematoceran Diptera. Midges are found on practically every land area outside permanently arid deserts and the frigid zones. Some midges, such as many Phlebotominae and Simuliidae, are vectors of various diseases. Many others play useful roles as prey for insectivores, such as various frogs and swallows. Others are important as detritivores, and form part of various nutrient cycles. The habits of midges vary greatly from species to species, though within any particular family, midges commonly have similar ecological roles.
Osmia cornifrons, also known as the horned-face bee, is a species of solitary bee indigenous to Northern Asia. Physically, this species of bee is recognized for its horn-like extensions originating from its lower face. Populations of O. cornifrons have been recorded in multiple locations, including Japan, Korea, China, and Russia. O. cornifrons are more docile as compared to other species of bees and are less prone to sting when aggravated.
Ceratopogonidae is a family of flies commonly known as no-see-ums, or biting midges, generally 1–3 millimetres in length. The family includes more than 5,000 species, distributed worldwide, apart from the Antarctic and the Arctic.
Phytophthora megakarya is an oomycete plant pathogen that causes black pod disease in cocoa trees in west and central Africa. This pathogen can cause detrimental loss of yield in the economically important cocoa industry, worth approximately $70 billion annually. It can damage any part of the tree, causing total yield losses which can easily reach 20-25%. A mixture of chemical and cultural controls, as well as choosing resistant plant varieties, are often necessary to control this pathogen.
Cacao swollen shoot virus (CSSV) is a plant pathogenic virus of the family Caulimoviridae that primarily infects cacao trees. It decreases cacao yield within the first year of infection, and usually kills the tree within a few years. Symptoms vary by strain, but leaf discoloration, stem/root swelling, and die-back generally occur. The virus is transmitted from tree to tree by mealybug vectors. It was first discovered in Ghana in 1936, and is currently endemic in Togo, Ghana and Nigeria. Over 200 million trees have already been claimed by this disease, which has prompted Ghana to launch the most ambitious and costly eradication effort of any country in the world against a viral plant disease.
The alkali bee, Nomia melanderi, is a ground-nesting bee native to deserts and semi-arid desert basins of the western United States. It was described by Theodore Dru Alison Cockerell in 1906. While solitary, these bees nest near each other and can form extremely dense aggregations in areas with favorable conditions.
The environmental impact of cocoa production includes deforestation, soil contamination, and herbicide resistance. The majority of cocoa farms are now located in Côte d'Ivoire and Ghana.
Thonningia is a monotypic genus of flowering plants in the family Balanophoraceae containing the single species Thonningia sanguinea. It is distributed throughout much of southern and western Africa, particularly the tropical regions. Common names for the plant include ground pineapple. A familiar plant to humans, it has an extremely long list of common names in many African languages. Many names are inspired by the resemblance of the plant's inflorescence to a pineapple or palm tree. Some of the names can be translated as pineapple of the bush, duiker's kolanut, and crown of the ground.
Blastophaga psenes is a wasp species in the genus Blastophaga. It pollinates the common fig Ficus carica and the closely related Ficus palmata. Without a colony or nest, these wasps breed in figs and the adults live for only a few days or weeks. They locate the fig they wish to pollinate through olfactory senses.
Black pod disease is a protozoal disease of Cocoa trees. This pathogen if left untreated can destroy all yields; annually the pathogen can cause a yield loss of up to 1/3 and up to 10% of total trees can be lost completely.
Pollination traps or trap-flowers are plant flower structures that aid the trapping of insects, mainly flies, so as to enhance their effectiveness in pollination. The structures of pollination traps can include deep tubular corollas with downward pointing hairs, slippery surfaces, adhesive liquid, attractants, flower closing and other mechanisms.
A gnat is any of many species of tiny flying insects in the dipterid suborder Nematocera, especially those in the families Mycetophilidae, Anisopodidae and Sciaridae. They can be both biting and non-biting. Most often they fly in large numbers, called clouds. "Gnat" is a loose descriptive category rather than a phylogenetic or other technical term, so there is no scientific consensus on what constitutes a gnat. Some entomologists consider only non-biting flies to be gnats. Certain universities and institutes also distinguish eye gnats: the Smithsonian Institution describes them as "non-biting flies, no bigger than a few grains of salt, ... attracted to fluids secreted by your eyes".
Cocoa necrosis virus (CoNV) is a plant pathogenic virus of the genus nepovirus that infects Theobroma cacao en natura causing cacao necrosis disease. CoNV is considered synonymous with Strain S of cacao swollen shoot virus. Unlike Cacao swollen shoot virus, it is not transmitted by mealybugs nor vectored by aphids, beetles, or leafhoppers that also commonly infest cacao. It is serologically, distantly related to Tomato black ring virus and very distantly related to Grapevine chrome mosaic virus.
Forcipomyia is a genus of biting midges in the subfamily Forcipomyiinae. Species of the subgenus Lasiohelea suck vertebrate blood. Some species are ectoparasites on larger insects. Other species in the genus are important pollinators of the cacao tree. There are at least 1,000 described species in Forcipomyia.
Theobroma speciosum is an arboriform species of flowering plant in the mallow family native to northern South America. It is the 35th most abundant species of tree in the Amazon rainforest.