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The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families, with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae.
Weevils are beetles belonging to the superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than 6 mm in length, and herbivorous. Approximately 97,000 species of weevils are known. They belong to several families, with most of them in the family Curculionidae. It also includes bark beetles, which while morphologically dissimilar to other weevils in lacking the distinctive snout, is a subfamily of Curculionidae. Some other beetles, although not closely related, bear the name "weevil", such as the biscuit weevil, which belongs to the family Ptinidae.
The New York weevil is a species of primitive weevil; large for weevils (12–18 mm), it is covered with fine bristles and has a regular pattern of light and dark spots. It occurs in the eastern United States and southern Canada.
The bean weevils or seed beetles are a subfamily (Bruchinae) of beetles, now placed in the family Chrysomelidae, though they have historically been treated as a separate family. They are granivores, and typically infest various kinds of seeds or beans, living most of their lives inside a single seed. The subfamily includes about 1,650 species and are found worldwide.
The beetle subfamily Curculioninae is part of the weevil family Curculionidae. It contains over 23,500 described species in 2,200 genera, and is therefore the largest weevil subfamily. Given that the beetle order (Coleoptera) contains about one-quarter of all known organisms, the Curculioninae represent one of the – if not the – most successful radiations of terrestrial Metazoa.
Dryophthorinae is a weevil subfamily within the family Curculionidae. While it is not universally accepted as distinct from other curculionid subfamilies, at least one major recent revision elevated it to family rank, as Dryophthoridae
The tribe Rhynchophorini is the largest member of the true weevil subfamily Dryophthorinae. Alonso-Zarazaga and Lyal (1999) treated it as a distinct subfamily, Rhynchophorinae. Weevils of this tribe have the pygidium not covered by the elytra.
Belidae is a family of weevils, called belids or primitive weevils because they have straight antennae, unlike the "true weevils" or Curculionidae which have geniculate (elbowed) antennae. They are sometimes known as "cycad weevils", but this properly refers to a few species from the genera Parallocorynus and Rhopalotria.
Brentidae is a cosmopolitan family of primarily xylophagous beetles also known as straight-snouted weevils. The concept of this family has been recently expanded with the inclusion of three groups formerly placed in the Curculionidae; the subfamilies Apioninae, Cyladinae, and Nanophyinae, as well as the Ithycerinae, previously considered a separate family. They are most diverse in the tropics, but occur throughout the temperate regions of the world. They are among the families of weevils that have non-elbowed antennae, and tend to be elongate and flattened, though there are numerous exceptions.
Anthribidae is a family of beetles also known as fungus weevils. The antennae are not elbowed, may occasionally be longer than the body and thread-like, and can be the longest of any members of Curculionoidea. As in the Nemonychidae, the labrum appears as a separate segment to the clypeus, and the maxillary palps are long and projecting.
Cryptorhynchinae is a large subfamily of weevils (Curculionidae), with some 6000 species. They are found in most zoogeographic regions although they are most diverse in the Neotropics, Australia and Oceania.
Lixinae is a subfamily of true weevils, included in the Molytinae in many older treatments. They are mainly root feeders, although some develop in flower buds or stems. Several species are used in biological control of invasive weeds, namely knapweeds (Centaurea).
The Attelabidae is a widespread family of weevils. They are among the primitive weevils, because of their straight antennae, which are inserted near the base of the rostrum. The prothorax is much narrower than the base of the elytra on the abdomen. Attelabidae and the related family Rhynchitidae are known commonly as the leaf-rolling weevils. Rhynchitidae may be treated as subfamily Rhynchitinae of the Attelabidae.
Nemonychidae is a small family of weevils, placed within the primitive weevil group because they have straight rather than geniculate (elbowed) antennae. They are often called pine flower weevils. As in the Anthribidae, the labrum appears as a separate segment to the clypeus, and the maxillary palps are long and projecting. Nemonychidae have all ventrites free, while Anthribidae have ventrites 1-4 connate or partially fused. Nemonychidae lack lateral carinae on the pronotum, while these are usually present, though may be short, in Anthribidae.
The tooth-nosed snout weevils, Rhynchitidae, are small beetles that are usually found in vegetation. They usually use buds, fruits, or seeds for oviposition. The tooth-nosed snout weevils receive this name due to the teeth on the edges of their mandibles.
The Entiminae are a large subfamily in the weevil family Curculionidae, containing most of the short-nosed weevils, including such genera as Entimus, Otiorhynchus, Phyllobius, Sitona, and Pachyrrhynchus. In comparison with their stunning diversity, only a few of these weevils are notorious pests of major economic importance. Entimines are commonly encountered in the field, including urban environments, and abundant in entomological collections.
Baridinae is a subfamily of true weevils (Curculionidae). It was established by Carl Johan Schönherr in 1836. Some 4,300 species in 550 genera are placed here, most of which occur in the New World. A few are economically significant pests, while others are in turn used for biocontrol of invasive plant pests. This subfamily also contains a few endangered species.
Polydrusini is a weevil tribe in the subfamily Entiminae.
Anthribinae is a subfamily of fungus weevils in the family of beetles known as Anthribidae. There are over 50 genera and more than 80 described species in Anthribinae.
Conoderinae is a subfamily of true weevils in the beetle family Curculionidae. There are more than 210 genera in 15 tribes, and about 2,400 described species in Conoderinae.
References
- Van Dyke, E.C. 1936: New species of North American Weevils in the family Curculionidae, subfamily Brachyrhininae, IV. The Pan-Pacific entomologist, 12(1): 19–32.
- Alonso-Zarazaga, M.A.; Lyal, C.H.C. 1999: A world catalogue of families and genera of Curculionoidea (Insecta: Coleoptera) (excepting Scolytidae and Platypodidae). Entomopraxis, Barcelona
