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Septa are thin walls or partitions between the internal chambers (camerae) of the shell of a cephalopod, namely nautiloids or ammonoids.
Discosorida are an order of cephalopods that lived from the beginning of the Middle Ordovician, through the Silurian, and into the Devonian. Discosorids are unique in the structure and formation of the siphuncle, the tube that runs through and connects the camerae (chambers) in cephalopods, which unlike those in other orders is zoned longitudinally along the segments rather than laterally. Siphuncle structure indicated that the Discosorida evolved directly from the Plectronoceratida rather than through the more developed Ellesmerocerida, as did the other orders. Finally and most diagnostic, discosorids developed a reinforcing, grommet-like structure in the septal opening of the siphuncle known as the bullette, formed by a thickening of the connecting ring as it draped around the folded back septal neck.
Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.
The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.
Clarkoceras is a genus of breviconic ellesmerocerid cephalopods, one of only two genera known to have crossed from the Late Cambrian, Trempealeauan, into the Early Ordovician, Gasconadian. ; the other being Ectenolites.
The Dresbachian is a Maentwrogian regional stage of North America, lasting from 501 to 497 million years ago. It is part of the Upper Cambrian and is defined by four trilobite zones. It overlaps with the ICS-stages Guzhangian, Paibian and the lowest Jiangshanian.
The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic).
Quebecoceras is an extinct genus from the nautiloid order Ellesmerocerida that lived during the Gasconadian Stage at the beginning of the Early Ordovician.
Basslerocerida is an order of nautiloid cephalopods from the Ordovician comprising exogastric longiconic cyrtocones, that is no longer in common use.
The Bassleroceratidae is a family of gradually expanding, smooth ellesmerocerids with a slight to moderate exogastric curvature, subcircular to strongly compressed cross section, and ventral orthochaonitc siphuncle. The ventral side is typically more sharply rounded than the dorsal side and septa are close spaced. Connecting rings are thick and slightly expanded into the siphuncle, making the segments slightly concave; characteristic of the Ellesmerocerida.
Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.
Balticoceras is a genus in the Orthocerid family, Orthoceratidae, from the Upper Ordovician of Estonia, closely related to Michelinoceras. Balticoceras is distinguished by its straight shells with a subcircular cross section, broadly flattened ventrally and evenly rounded dorsally, and by its marginal, orthochoanitic siphuncle. Balticoceras which is included in the subfamily Michelinoceratinae along with Michelinoceras should not be confused with Baltoceras which belongs in the Ellesmerocerida.
Bassleroceras is an elongate upwardly curved, exogastric, genus with the venter on the under side more sharply rounded than the dorsum on the upper. The siphuncle is ventral, composed of thick-walled tubular segments in which connection rings thicken in towardly as in both the Ellesmerocerida and primitive Tarphycerida.
Dakeoceras is a genus of simple cyrtoconic ellesmeroceratid cephalopods in the fossil record. All known species come from the Lower Canadian epoch (Gasconadian) of North America.
Burenoceras is a genus of small, even tiny, Gasconadian cyrtoconic ellesmeroceratids in which the phragmocone—the chambered portion—is shorter that the living chamber.
Plectronoceratoidea is a superorder or subclass containing primitive nautiloids from the Late Cambrian and Early Ordovician. This group is best considered a paraphyletic grade of early cephalopods, as it contains the ancestors of subsequent post-Cambrian cephalopod orders.
Ellesmeroceras is the type genus for the Ellesmeroceratidae, a family of primitive nautiloid cephalopods, that is characterized by its small, generally compressed, gradually expanded, orthoconic shell, found in Lower Ordovician marine sediments. The septa are close spaced and the siphuncle is ventral, about 0.2 the diameter of the shell. Septal necks are typically orthochoanitic but may slant inwardly (loxochoanitic) or reach halfway to the previous septum (hemichoanitic). Connecting rings are thick. As common for the Ellesmerocerida, Ellesmeroceras has diaphragms within the siphuncle tube.
Albertoceras is a genus of Early Ordovician ellesmeroceratids with a small, slender, orthoconic to slightly endogastric shell; some even tiny. The cross section is strongly compressed so as to make the height proportionally notably greater than the width. The sutures, marking the edges of the septa, have broad, shallow, lateral lobes. The siphuncle, which seems to lack diaphragms, lies close to the venter.
Yanhecerida is a small order of Late Cambrian nautiloid cephalopods. They were similar to the more diverse Plectronocerida and Ellesmerocerida, with short shells, closely-spaced septa, and diaphragms (partitions) within the siphuncle. Their most distinctive trait is the conical shape of the diaphragms, similar to the endocones which characterize the later nautiloid order Endocerida. While some authors have argued that Yanhecerida should be lumped into Ellesmerocerida, a phylogenetic analysis has maintained that Yanhecerida is a valid clade of early cephalopods closely related to a paraphyletic Ellesmerocerida.
Multiceratoidea is a major subclass or superorder of Paleozoic nautiloid cephalopods. Members of this group can be characterized by nautilosiphonate connecting rings, with an organic inner layer and outer layer of calcitic spherules and blades, similar to the modern nautilus. The earliest-diverging multiceratoids have oncomyarian muscle scars, though several orders trend towards a ventromyarian condition. Multiceratoid shells are generally short and curled, with a relatively small aperture (opening). Cameral deposits are never found among the multiceratoids, though several orders are known to bear endosiphuncular deposits within their siphuncles.
References
 | This article includes a list of references, related reading, or external links, but its sources remain unclear because it lacks inline citations .(March 2015) |
- W.M.Furnish & Brian F. Glenister, 1964. Nautiloidea - Ellesmerocerida; Treatise on Invertebrate Paleontology, Part K. Geological Society of America.
- Lawrenceoceras in Fossilworks
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