Bassleroceratidae

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Bassleroceratidae
Temporal range: Lower Ordovician
Scientific classification
Kingdom: Animalia
Phylum: Mollusca
Class: Cephalopoda
Subclass: Nautiloidea
Order: Ellesmerocerida
Family:Bassleroceratidae
Ulrich et al (1944)

The Bassleroceratidae is a family of gradually expanding, smooth ellesmerocerids with a slight to moderate exogastric curvature, subcircular to strongly compressed cross section, and ventral orthochaonitc siphuncle. The ventral side is typically more sharply rounded than the dorsal side and septa are close spaced. Connecting rings are thick and slightly expanded into the siphuncle, making the segments slightly concave; characteristic of the Ellesmerocerida. [1]

The Ellesmerocerida is an order of primitive cephalopods belonging to the subclass Nautiloidea with a widespread distribution that lived during the Late Cambrian and Ordovician.

Basslerocerids are limited to the Lower Ordovician [1] and first appeared sometime in the Gasconadian, (Tremedocian) They gave rise, possibly through Bassleroceras, by evolving and ever tightened curvature to the Tarphycerida [2] and by a thinning of the connecting rings to the Graciloceratidae which are ancestral Oncocerida. [3] Furthermore, they may have given rise through some form like Bassleroceras or Lawrenceoceras to directly to the Oncoceratid genus Richardsonoceras.

The Tarphycerida were the first of the coiled cephalopods, found in marine sediments from the Lower Ordovician to the Middle Devonian. Some, such as Aphetoceras and Estonioceras, are loosely coiled and gyroconic; others, such as Campbelloceras, Tarphyceras, and Trocholites, are tightly coiled, but evolute with all whorls showing. The body chamber of tarphycerids is typically long and tubular, as much as half the length of the containing whorl in most, greater than in the Silurian Ophidioceratidae. The Tarphycerida evolved from the elongated, compressed, exogastric Bassleroceratidae, probably Bassleroceras, around the end of the Gasconadian through forms like Aphetoceras. Close coiling developed rather quickly, and both gyroconic and evolute forms are found in the early middle Canadian.

Oncocerida order of molluscs

The Oncocerida comprise a diverse group of generally small nautiloid cephalopods known from the Middle Ordovician to the Mississippian, in which the connecting rings are thin and siphuncle segments are variably expanded. At present the order consists of some 16 families, a few of which, such as the Oncoceratidae, Brevicoceratidae, and Acleistoceratidae contain a fair number of genera each while others like the Trimeroceratidae and Archiacoceratidae are represented by only two or three.

The Bassleroceratidae was named by Ulrich et al., 1944 [1] and assigned to the Basslerocerida, an order proposed by Flower (1950) intermediary between the Ellesmerocerida and the Tarphycerida, which also included the Graciloceratidae. Flower later abandoned the Basslerocerida and added the Bassleroceratidae to its descendant group, the Tarphycerida. Furnish and Glenister on the other hand included the Bassleroceratidae in with its ancestral group, the Ellesmerocerida, where it is generally assigned.

The Graciloceratidae is a family of nautiloid cephalopods from the Middle and Upper Ordovician belonging to the Oncocerida, characterized by exogastric cyrtocones that expand slightly or moderately and have thin walled, orthochoantic marginal or subventral, tubular siphuncles.

Related Research Articles

Plectronocerida order of molluscs

Plectronocerida is a primitive order from which subsequent cephalopod orders are ultimately derived.

Trocholites is a tarphycerid genus in the family Trocholitidae from the Middle and Upper Ordovician with a gradually expanding, weakly ribbed shell; whorls in contact, dorsum slightly impressed; cross section depressed, venter and sides rounded; siphuncle close to but not at the dorsal margin.

The Ellesmeroceratidae constitute a family within the cephalopod order Ellesmerocerida. They lived from the Upper Cambrian to the Lower Ordovician. They are characterized by straight and endogastric shells, often laterally compressed, so the dorso-ventral dimension is slightly greater than the lateral, with close spaced sutures having shallow lateral lobes and a generally large tubular ventro-marginal siphuncle with concave segments and irregularly spaced diaphragms. Connecting rings are thick and layered, externally straight but thickening inwardly with the maximum near the middle of the segment so as to leave concave depressions on internal siphuncle molds. Septal necks are typically orthochoanitic but vary in length from almost absent (achoanitic) to reaching halfway to the previous septum (hemichoanitic) and may even slope inwardly (loxochoanitic).

The Tarphyceratidae are tightly coiled, evolute Tarphycerida with ventral siphuncles. The dorsum is characteristically impressed where the whorl presses against the venter of the previous. The Tarphyceratidae are derived from Bassleroceras or possibly from some member of the Estonioceratidae.

Baltoceratidae is an extinct family of orthoconic cephalopods belonging to the subclass Nautiloidea endemic to what would be Asia, Australia, Europe, North America, and South America during the Ordovician living from about 480–460 mya, existing for approximately 20 million years.

Basslerocerida is an order of nautiloid cephalopods from the Ordovician comprising exogastric longiconic cyrtocones, that is no longer in common use.

Oncoceratidae family of molluscs

Oncoceratidae is a family of nauatiloid cephalopods in the order Oncocerida established by Hyatt, 1884, that range from the Middle Ordovician to the Upper Silurian.

Protcycloceratidae is an extinct family of slender, commonly annulate, members of the cephalopod order Ellesmerocerida that lived during the Early Ordovician.

The Lituitidae are a family of evolved tarphycerids characterized by a long orthoconic section that follows a coiled juvenile portion at the apex, along with a generally tubular siphuncle, which like that of the barrandeocerids is composed of thin connecting rings.

Bassleroceras is an elongate upwardly curved, exogastric, genus with the venter on the under side more sharply rounded than the dorsum on the upper. The siphuncle is ventral, composed of thick-walled tubular segments in which connection rings thicken in towardly as in both the Ellesmerocerida and primitive Tarphycerida.

Estonioceratidae family of molluscs

The Estonioceratidae are a family of loosely coiled tarphycerids in which the inner side of the whorls, which forms the dorsum, is rounded or flat with no impression, and in which the siphuncle, composed of thick tubular segments, is located ventrally. The Estonioceratidae seem to form a link between the ancestral Bassleroceratidae and the more tightly coiled Tarphyceratidae.

The Nautilitoidea is a superorder within the subclass Nautiloidea, comprising the phylogenetically related Nautilida, Oncocerida, and Tarphycerida.

Ellesmeroceras is the type genus for the Ellesmeroceratidae, a family of primitive nautiloid cephalopods, that is characterized by its small, generally compressed, gradually expanded, orthoconic shell, found in Lower Ordovician marine sediments. The septa are close spaced and the siphuncle is ventral, about 0.2 the diameter of the shell. Septal necks are typically orthochoanitic but may slant inwardly (loxochoanitic) or reach halfway to the previous septum (hemichoanitic). Connecting rings are thick. As common for the Ellesmerocerida, Ellesmeroceras has diaphragms within the siphuncle tube.

Alaskoceras is a genus of lower Ordovician coiled nautiloid cephalopods; the shell moderately expanded, ribbed, with a divergent living chamber; whorl section more broadly rounded ventrally than dorsally; siphuncle marginal at maturity, septal necks short, almost achoanitic; connecting rings thick, layered.

Leonardoceras is a genus of nautiloid cephalopods from the lower Middle Ordovician of Nevada (USA), the shell a small slender exogastric cyrtocone with the venter more narrowly rounded than the dorsum, resembling in overall form a small Bassleroceras. Septa are close spaced, the living chamber short, the siphuncle close to the venter.

Lawrenceoceras is a genus of moderately curved, gently expanding bassleroceratids from the Lower Ordovician of eastern North America. Septa dividing the chambers are close spaced; sutures straight, transverse; the siphuncle narrow, submarginal.

References

  1. 1 2 3 Furnish and Glenister 1964a, Nautiloidea-Ellesmerocerida; Treatise on Invertebrate Paleontology, Part K Nautiloidea.
  2. Furnish and Glenister 1964b, Nautiloidea-Tarphycerida; Treatise on Invertebrate Paleontology Part K Nautiloidea.
  3. Sweet 1964, Nautiloidea-Oncocerida; Treatise on Invertebrate Paleontology Part K Nautiloidea.