Chalcid wasps are insects within the superfamily Chalcidoidea, part of the order Hymenoptera. The superfamily contains some 22,500 known species, and an estimated total diversity of more than 500,000 species, meaning the vast majority have yet to be discovered and described. The name "chalcid" is often confused with the name "chalcidid", though the latter refers strictly to one constituent family, the Chalcididae, rather than the superfamily as a whole; accordingly, most recent publications (e.g.,) use the name "chalcidoid" when referring to members of the superfamily.
In evolutionary ecology, a parasitoid is an organism that lives in close association with its host at the host's expense, eventually resulting in the death of the host. Parasitoidism is one of six major evolutionary strategies within parasitism, distinguished by the fatal prognosis for the host, which makes the strategy close to predation.
Kleptoparasitism is a form of feeding in which one animal deliberately takes food from another. The strategy is evolutionarily stable when stealing is less costly than direct feeding, such as when food is scarce or when victims are abundant. Many kleptoparasites are arthropods, especially bees and wasps, but including some true flies, dung beetles, bugs, and spiders. Cuckoo bees are specialized kleptoparasites which lay their eggs either on the pollen masses made by other bees, or on the insect hosts of parasitoid wasps. They are an instance of Emery's rule, which states that insect social parasites tend to be closely related to their hosts. The behavior occurs, too, in vertebrates including birds such as skuas, which persistently chase other seabirds until they disgorge their food, and carnivorous mammals such as spotted hyenas and lions. Other species opportunistically indulge in kleptoparasitism.
Apocrita is a suborder of insects in the order Hymenoptera. It includes wasps, bees, and ants, and consists of many families. It contains the most advanced hymenopterans and is distinguished from Symphyta by the narrow "waist" (petiole) formed between the first two segments of the actual abdomen; the first abdominal segment is fused to the thorax, and is called the propodeum. Therefore, it is general practice, when discussing the body of an apocritan in a technical sense, to refer to the mesosoma and metasoma rather than the "thorax" and "abdomen", respectively. The evolution of a constricted waist was an important adaption for the parasitoid lifestyle of the ancestral apocritan, allowing more maneuverability of the female's ovipositor. The ovipositor either extends freely or is retracted, and may be developed into a stinger for both defense and paralyzing prey. Larvae are legless and blind, and either feed inside a host or in a nest cell provisioned by their mothers.
The Ichneumonidae, also known as ichneumon wasps, ichneumonid wasps, ichneumonids, or Darwin wasps, are a family of parasitoid wasps of the insect order Hymenoptera. They are one of the most diverse groups within the Hymenoptera with roughly 25,000 species described as of 2016. However, this likely represents less than a quarter of their true richness as reliable estimates are lacking, along with much of the most basic knowledge about their ecology, distribution, and evolution. It is estimated that there are more species in this family than there are species of birds and mammals combined. Ichneumonid wasps, with very few exceptions, attack the immature stages of holometabolous insects and spiders, eventually killing their hosts. They play an important role as regulators of insect populations, both in natural and semi-natural systems, making them promising agents for biological control.
The organ pipe mud dauber is a predatory wasp in the family Crabronidae. It is fairly large, ranging from 3.9–5.1 cm, and has been recorded to fly from May to September. Females and males are similar in colour, a shiny black, with the end part of the back leg being pale yellow to white. The organ pipe mud dauber feeds mainly on three genera of spider: Neoscona, Araneus, and Eustala. Melittobia, a parasitoid wasp, is a common ectoparasite of T. politum prepupae. Other sources of parasitism include the bombyliid fly Anthrax, chrysidid wasps, and various species of scavenger flies (Miltogramminae). The tufted titmouse is a known predator of T. politum, and may feed on them more commonly than previously thought, as the holes made by the titmouse are similar in shape and size to those made by T. politum leaving the nest after pupation.
The Eucharitidae are a family of parasitic wasps. Eucharitid wasps are members of the superfamily Chalcidoidea and consist of four subfamilies: Akapalinae, Eucharitinae, Gollumiellinae, and Oraseminae. Most of the 42 genera and >400 species of Eucharitidae are members of the subfamilies Oraseminae and Eucharitinae, and are found in tropical regions of the world.
The Pteromalidae are a large family of wasps, the majority being parasitoids of other insects. They are found throughout the world in virtually all habitats, and many are important as biological control agents. The oldest known fossil is known from the Early Cretaceous.
The Leucospidae are a specialized group of wasps within the superfamily Chalcidoidea, that are ectoparasitoids of aculeate wasps or bees. They are typically mimics of bees or stinging wasps, often black with yellow, red, or white markings, sometimes metallic, with a robust mesosoma and very strong sculpturing. The hind femora are often greatly enlarged, with a row of teeth or serrations along the lower margin as in Chalcididae. The wing has a longitudinal fold. The female ovipositor is sometimes short, but if not, it is recurved and lies along the dorsal side of the metasoma, a unique feature. The males are also unusual, in the fusion of many of the metasomal segments to form a capsule-like "carapace".
Nasonia are a genus of small pteromalid parasitoid wasps that sting and lay eggs in the pupae of various flies. The fly species that Nasonia usually parasitize are primarily blow flies and flesh flies, making Nasonia a useful tool for biocontrol of these pest flies. The small match-head sized wasps are also referred to as jewel wasps based on the emerald sheen of their exoskeleton.
The European paper wasp is one of the most common and well-known species of social wasps in the genus Polistes. Its diet is more diverse than those of most Polistes species—many genera of insects versus mainly caterpillars in other Polistes—giving it superior survivability compared to other wasp species during a shortage of resources.
Nasonia vitripennis is one of four known species under the genus Nasonia - small parasitoid wasps that afflict the larvae of parasitic carrion flies such as blowflies and flesh flies, which themselves are parasitic toward nestling birds. It is the best known and most widely studied of the parasitoid wasps, and their study forms a vital part of the information used to describe the order Hymenoptera, along with information from bees and ants. This parasitoid behaviour makes the wasps an interest for the development of biopesticide and biological systems for controlling unwanted insects.
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. The wasps do not constitute a clade, a complete natural group with a single ancestor, as bees and ants are deeply nested within the wasps, having evolved from wasp ancestors. Wasps that are members of the clade Aculeata can sting their prey.
Polistes metricus is a wasp native to North America. In the United States, it ranges throughout the southern Midwest, the South, and as far northeast as New York, but has recently been spotted in southwest Ontario. A single female specimen has also been reported from Dryden, Maine. P. metricus is dark colored, with yellow tarsi and black tibia. Nests of P. metricus can be found attached to the sides of buildings, trees, and shrubbery.
Nasonia giraulti is a species of pteromalid wasp in the family Pteromalidae. It can be differentiated from other species in its genus by its antennae. The species can be found in eastern North America. It is a parasitoid of bird blow fly pupae.
Polistes exclamans, the Guinea paper wasp, is a social wasp and is part of the family Vespidae of the order Hymenoptera. It has been found in Ontario, Canada and the eastern United States from Illinois down south to Florida and west to Nebraska and California. It is also found in Mexico from Chihuahua to Jalisco, Hidalgo. P. exclamans has shown variability in its range including an absence of the species in eastern Missouri from the 1920’s to 1940’s, a presence in the 1960’s to 1980’s, and an apparent absence again of the species in these same sites in eastern Missouri since 1989. This suggests that their range has either expanded northward and contracted southward or that they have large, long-term cycles of abundance. P. exclamans has three specific castes, including males, workers, and queens, but the dominance hierarchy is further distinguished by age. The older the wasp is, the higher it is in ranking within the colony. In most P. exclamans nests, there is one queen who lays all the eggs in the colony. The physiological similarities between the worker and queen castes have led to experiments attempting to distinguish the characteristics of these two castes and how they are determined, though males have easily identifiable physiological characteristics. Since P. exclamans live in relatively small, open combed nests, they are often subject to predators and parasites, such as Chalcoela iphitalis, Elasmus polistis, and birds. P. exclamans have defense and recognition strategies that help protect against these predators and parasites.
Arsenophonus nasoniae is a species of bacterium which was previously isolated from Nasonia vitripennis, a species of parasitoid wasp. These wasps are generalists which afflict the larvae of parasitic carrion flies such as blowflies, houseflies and flesh flies. A. nasoniae belongs to the phylum Pseudomonadota and family Morganellaceae. The genus Arsenophonus, has a close relationship to the Proteus (bacterium) rather than to that of Salmonella and Escherichia. The genus is composed of gammaproteobacterial, secondary-endosymbionts which are gram-negative. Cells are non-flagellated, non-motile, non-spore forming and form long to highly filamentous rods. Cellular division is exhibited through septation. The name 'Arsenophonus nasoniae gen. nov., sp. nov.' was therefore proposed for the discovered bacterium due to its characteristics and its microbial interaction with N. vitripennis. The type strain of A. nasoniae is Strain SKI4.
Arsenophonus is a genus of Morganellaceae, of the Gammaproteobacteria. Members of the Arsenophonus genus are increasingly discovered bacterial symbionts of arthropods that are estimated to infect over 5% of arthropod species globally and form a variety of relationships with hosts across the mutualism parasitism continuum. Arsenophonus bacteria have been identified in a diversity of insect taxa, including economically important species such as the Western honey bee and the rice pest Nilaparvata lugens.
Polistes nimpha is a eusocial paper wasp found all over Europe, with particular sightings in Turkey, Finland, Estonia, and Latvia. It is also found in northern Africa, Pakistan, Iran, India, Kazakhstan, Mongolia, and China. The climate in these areas is relatively cold and snowy in the winter, while summers are usually hot and dry, with steppe vegetation. Polistes nimpha colonies are relatively small and easily manipulated.
Chlorocytus is a genus of wasps belonging to the family Pteromalidae.
