| Obtectomera | |
|---|---|
 | |
| Pyrausta purpuralis | |
Scientific classification  | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Class: | Insecta |
| Order: | Lepidoptera |
| Clade: | Neolepidoptera |
| Infraorder: | Heteroneura |
| Clade: | Eulepidoptera |
| Clade: | Ditrysia |
| Clade: | Apoditrysia |
| Clade: | Obtectomera Minet, 1986 |
The Obtectomera is a clade of macro-moths and butterflies, comprising over 100,000 species in at least 12 superfamilies. [1] [2]
The Obtectomera includes the following 12 superfamilies: [1]
Some other superfamilies are sometimes included: [2]
The macroheteroceran superfamilies were previously place in the Macrolepidoptera, but recent molecular studies have failed to recover the Macrolepidoptera as a monophyletic group. [1] [2] The latter grouping also included true butterflies (Papilionoidea), New World butterfly-moths (Hedylidae), Old World butterfly-moths (Calliduloidea), and European gold moths (Axioidea).
The Psychidae are a family of the Lepidoptera. The bagworm family is fairly small, with about 1,350 species described. Bagworm species are found globally, with some, such as the snailcase bagworm, in modern times settling continents where they are not native.
Macrolepidoptera is a group within the insect order Lepidoptera. Traditionally used for the larger butterflies and moths as opposed to the "microlepidoptera", this group is artificial. However, it seems that by moving some taxa about, a monophyletic macrolepidoptera can be easily achieved. The two superfamilies Geometroidea and Noctuoidea account for roughly one-quarter of all known Lepidoptera.
Riodinidae is the family of metalmark butterflies. The common name "metalmarks" refers to the small, metallic-looking spots commonly found on their wings. The 1,532 species are placed in 146 genera. Although mostly Neotropical in distribution, the family is also represented both in the Nearctic, Palearctic, Australasian (Dicallaneura), Afrotropic, and Indomalayan realms.
Nolidae is a family of moths with about 1,700 described species worldwide. They are mostly small with dull coloration, the main distinguishing feature being a silk cocoon with a vertical exit slit. The group is sometimes known as tuft moths, after the tufts of raised scales on the forewings of two subfamilies, Nolinae and Collomeninae. The larvae also tend to have muted colors and tufts of short hairs.
Sesioidea is a superfamily containing clearwing moths (Sesiidae), castniid moths (Castniidae) and little bear moths (Brachodidae). There is evidence from head and thoracic morphology that the first two families, internally feeding in plants as caterpillars, are sisters, whilst some brachodids are known to feed on leaf surfaces. Sesioidea is closely related to Cossoidea, which contains the also internal-feeding Goat and Leopard moths, and recent taxonomic treatments consider the sessoid families as part of Cossoidea sensu lato.
Mimallonidae Burmeister (mimallonids), sometimes known as "sack-bearer" moths for the larval case-building behavior, are a family of Lepidoptera containing over 300 named species in 43 genera. These moths are found only in the New World, with most taxa occurring in the Neotropics. Adult moths are externally similar to those belonging to some of the other Macroheterocera families Bombycoidea and Drepanoidea, and thus have been variously treated as belonging to either one of these or other superfamilies.
Gelechioidea is the superfamily of moths that contains the case-bearers, twirler moths, and relatives, also simply called curved-horn moths or gelechioid moths. It is a large and poorly understood '"micromoth" superfamily, constituting one of the basal lineages of the Ditrysia.
Cossoidea is the superfamily of moths that includes carpenter moths and relatives. Like their likely sister group Sesioidea they are internal feeders and have spiny pupae with moveable segments to allow them to extrude out of their exit holes in stems and trunks during emergence of the adult.
Aluctoidea is the superfamily of many-plumed and false plume moths. These small moths are most easily recognized by their wings. These each consist of many narrow strips of membrane around the major veins, instead of a continuous sheet of membrane between the veins. In living moths in the wild, this is often hard to see however. When they are at rest, the "plumes" partly overlap, appearing as solid wings. But even then, they can be recognized by the wings having a marked lengthwise pattern and uneven edge.
Hedylidae, the "American moth-butterflies", is a family of insects in the order Lepidoptera, representing the superfamily Hedyloidea. They have traditionally been viewed as an extant sister group of the butterfly superfamily Papilionoidea, but a 2014 phylogenetic analyses has suggested Hedylidae is a subgroup of Papilionoidea, and not a sister group, and are more accurately referred to as butterflies rather than moths. They are represented by a single Neotropical genus Macrosoma with 35 currently recognized species.
Carposinoidea, the "fruitworm moths", is a superfamily of insects in the lepidopteran order. The superfamily is also known as Copromorphoidea, which is a junior synonym. These moths are small to medium-sized and are broad-winged bearing some resemblance to the superfamilies Tortricoidea and Immoidea. The antennae are often "pectinate" especially in males, and many species of these well camouflaged moths bear raised tufts of scales on the wings and a specialised fringe of scales at the base of the hindwing sometimes in females only; there are a number of other structural characteristics. The position of this superfamily is not certain, but it has been placed in the natural group of "Apoditrysia" "Obtectomera", rather than with the superfamilies Alucitoidea or Epermenioidea within which it has sometimes previously been placed, on the grounds that shared larval and pupal characteristics of these groups have probably evolved independently. It has been suggested that the division into two families should be abandoned.
Acanthopteroctetidae is a small family of primitive moths with two described genera, Acanthopteroctetes and Catapterix, and a total of seven described species. They are known as the archaic sun moths.
Prodidactis mystica is an enigmatic Pyraloidea-like moth from southern Africa which has been placed in its own family, Prodidactidae and which belongs in the lepidopteran group Apoditrysia. Its closest relative amongst this large group of Lepidoptera is uncertain, but morphological and molecular evidence suggests placement of Prodidactidae in Hyblaeoidea. The larval food plant is Nuxia congesta (Stilbaceae).
Axia, the gold moths, is a genus of moths whose precise relationships within the macrolepidoptera are currently uncertain, but they currently are placed within the superfamily Drepanoidea. Uniquely, they have a pair of pocket-like organs on the seventh abdominal spiracle of the adult moth which are possibly sound receptive organs. They are quite large and brightly coloured moths that occur only in southern Europe and feed on species of Euphorbia. Sometimes they are attracted to light. The genus was first described by Jacob Hübner in 1821.
The insect order Lepidoptera consists of moths, most of which are night-flying, and a derived group, mainly day-flying, called butterflies. Within Lepidoptera as a whole, the groups listed below before Glossata contain a few basal families accounting for less than 200 species; the bulk of Lepidoptera are in the Glossata. Similarly, within the Glossata, there are a few basal groups listed first, with the bulk of species in the Heteroneura. Basal groups within Heteroneura cannot be defined with as much confidence, as there are still some disputes concerning the proper relations among these groups. At the family level, however, most groups are well defined, and the families are commonly used by hobbyists and scientists alike.
Cimeliidae, the gold moths, is a family of moths that is now placed in the macroheteroceran superfamily Drepanoidea, although previously placed in its own superfamily. Uniquely, they have a pair of pocket-like organs on the seventh abdominal spiracle of the adult moth which are only possibly sound receptive organs. They are quite large and brightly coloured moths that occur in southern Europe and feed on species of Euphorbia. Sometimes they are attracted to light. The family was first described by Pierre Chrétien in 1916.
Argyresthiidae is a family of moths known as the shiny head-standing moths. It was previously treated as a subfamily of Yponomeutidae.
Maliattha blandula is a species of moth in the family Noctuidae first described by Achille Guenée in 1862. It is found in Tanzania, Madagascar, Mauritius, Réunion and the Comoros. It was placed in the genus Maliattha in 2016.
Epimarptidae was a former, or is a possible, monotypic family of moths in the moth superfamily Gelechioidea. It can now be seen as either a synonym of family Batrachedridae, or a monotypic subfamily of that family.
The Macroheterocera are a well supported clade of moths that are closely related to butterflies and macro-moths.