Orcinus meyeri Temporal range: Early Miocene | |
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The holotype and only specimen | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Artiodactyla |
Infraorder: | Cetacea |
Family: | Delphinidae |
Genus: | Orcinus |
Species: | †O. meyeri |
Binomial name | |
†Orcinus meyeri (Brandt, 1873) | |
Synonyms | |
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Orcinus meyeri is a fossil species of Orcinus (killer whales) found in the Early Miocene deposits of southern Germany, known from two jaw fragments and 18 isolated teeth. It was originally described as Delphinus acutidens in 1859, but reclassified in 1873. Its validity is disputed, and it may be a synonymous with the ancient sperm whale Physeterula dubusi . It was found in the Alpine town of Stockach in the Molasse basin, which was a coastal area with strong tidal currents.
The remains of Orcinus meyeri was first described in 1859 by German paleontologist Christian Erich Hermann von Meyer as an ancient common dolphin, Delphinus acutidens, based on two jaw fragments and some isolated teeth found near the south German town of Stockach. [1] He also suggested the names Delphinus (Beluga) acutidens and D. (Orcopsis) acutidens.[ citation needed ] The species name acutidens derives from Latin acutus "sharp" and dens "teeth".[ citation needed ] In 1873, German naturalist Johann Friedrich von Brandt gave it the species name meyeri, citing "acutidens" as being inaccurate given the apparent bluntness of the teeth, and placed it into the same genus as the killer whale (Orcinus orca), which at the time was Orca, given the similarity of the teeth, reclassifying the whale as Orca meyeri. The remains were then kept in the Stuttgart Museum of Natural History. [2] In 1898, Swiss marine biologist Théophile Rudolphe Studer said O. meyeri was a subjective synonym of Delphinus acutidens, and there is room for debate on the question of the former's validity. [3] In 1904, French zoologist Édouard Louis Trouessart replaced Orca with Orcinus and described the whale as Orcinus meyeri. [4] In 1905, Austrian paleobiologist Othenio Abel considered D. acutidens to be synonymous with the sperm whale Physeterula dubusi. However, he was unsure if O. meyeri was also a synonym. [5]
The largest of the jaw fragments measured 26.2 cm (10.3 in) in length and 8 cm (3.1 in) in height. 18 isolated teeth were found, ranging in size from 5–6 cm (2.0–2.4 in) in height and 1.8–2 cm (0.71–0.79 in) in width at the base. [2] In comparison, the modern killer whale has teeth about 10–13 centimetres (4–5 in) in height and 2.5 cm (1 in) in diameter. [6] The animal in life would probably have had 48 conical teeth in total, [2] in comparison to the modern killer whale (O. orca) which has 40 to 56. [6] The teeth of O. meyeri are distinct from the modern killer whale by having two vertical grooves originating at the tip. [2]
The Orcinus lineage, like many other predatory marine lineages, may have fished up the food chain and progressively evolved to eat bigger and bigger food items, with Pliocene killer whales able to hunt large fish, and the modern killer whale able to hunt large whales. [7]
Stockach is situated in the Molasse basin, which dates to the Early Miocene, and was submerged in the Western Paratethys Sea. The basin represents coastal waters and strong tidal currents, with an average depth of less than 50 m (160 ft). [8] Central Europe, at this time, probably represented an upwelling area along a continental shelf, which attracted a variety of sea life, including swarming fish. [9] Land was probably dominated by nearshore swamps which emptied into the sea, and the area featured ancient beavers, hedgehogs, several river turtle species, and various other semi-aquatic creatures. [10] The sea progressively moved southwards, and the connection to the ocean closed about 17 million years ago (mya), turning the area into a system of brackish and freshwater lakes. [11]
The orca, also called killer whale, is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only extant species in the genus Orcinus and is recognizable by its black-and-white patterned body. A cosmopolitan species, orcas can be found in all of the world's oceans in a variety of marine environments, from Arctic and Antarctic regions to tropical seas.
The toothed whales are a parvorder of cetaceans that includes dolphins, porpoises, and all other whales possessing teeth, such as the beaked whales and sperm whales. 73 species of toothed whales are described. They are one of two living groups of cetaceans, the other being the baleen whales (Mysticeti), which have baleen instead of teeth. The two groups are thought to have diverged around 34 million years ago (mya).
Orcinus is a genus of Delphinidae, the family of carnivorous marine mammals known as dolphins. It includes the largest delphinid species, Orcinus orca, known as the orca or killer whale. Two extinct species are recognised, Orcinus paleorca and O. citoniensis, describing fossilised remains of the genus. The other extinct species O. meyeri is disputed.
Old Tom was a killer whale (orca) known to whalers in the port of Eden, New South Wales, on the southeast coast of Australia. Old Tom measured 22 feet (6.7 m) and weighed 6 tons, with a 3.33 ft (1.01 m) skull and teeth about 5.31 inches (135 mm) long.
The Cariaco Basin lies off the north central coast of Venezuela and forms the Gulf of Cariaco. It is bounded on the east by Margarita Island, Cubagua Island, and the Araya Peninsula; on the north by Tortuga Island and the Tortuga Banks; on the west by Cape Codera and the rocks known as Farallón Centinela; and on the south by the coast of Venezuela.
Zygophyseter varolai is an extinct sperm whale that lived during the Tortonian age of the Late Miocene 11.2 to 7.6 million years ago. It is known from a single specimen from the Pietra Leccese Formation in Italy. It was a member of a stem group of fossil macroraptorial sperm whales also including Brygmophyseter, Acrophyseter, and Livyatan. It probably grew to be around 6.5 to 7 meters in length and shared some characteristics with other raptorials, such as large teeth with tooth enamel that were functional in both the upper and lower jaws which the modern sperm whale lacks. It also had a beak, the ability to echolocate prey, and could have probably swum faster than the modern-day sperm whale which can reach 4 kilometers per hour (2.5 mph). These were probably used in the capture of large prey, such as large fish, seals, and whales. In fact, its common name, the killer sperm whale, refers to its feeding habits that would have had a resemblance to the modern-day killer whale.
Livyatan is an extinct genus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Miocene epoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5 mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.
Acrophyseter is a genus of extinct sperm whales that lived in the Late Miocene off the coast of Peru comprising two species: A. deinodon and A. robustus. It is part of a group of macroraptorial sperm whales which all shared several features for the purpose of hunting large prey, such as deeply-rooted and thick teeth. Acrophyseter measured 4–4.5 metres (13–15 ft), making it the smallest raptorial sperm whale. Because of its short pointed snout, and its strong curved front teeth, it probably fed on the large marine vertebrates of its time, such as seals and other whales.
Orcinus citoniensis is an extinct species of killer whale identified in the Late Pliocene of Italy and the Early Pleistocene of England. It was smaller than the modern killer whale, 4 m (13 ft) versus 7 to 10 m, and had around 8 more teeth in its jaw. It may have resembled the modern killer whale in appearance, and could represent a transitional species between the modern killer whale and other dolphins. O. citoniensis could have hunted fish and squid in pods, and coexisted with other large predators of the time such as the orcinine Hemisyntrachelus and the extinct shark Otodus megalodon.
Squalodontidae or the shark-toothed dolphins is an extinct family of large toothed whales who had long narrow jaws. Squalodontids are known from all continents except Antarctica, from the Oligocene to the Neogene, but they had a maximal diversity and global distribution during the Late Oligocene and Early to Middle Miocene.
Scaldicetus is an extinct genus of highly predatory macroraptorial sperm whale. Although widely used for a number of extinct physeterids with primitive dental morphology consisting of enameled teeth, Scaldicetus as generally recognized appears to be a wastebasket taxon filled with more-or-less unrelated primitive sperm whales.
Orcinus paleorca is a fossil species of Orcinus, the genus of killer whales (orca), dated to the Middle Pleistocene. The only known specimen is a tooth fragment discovered in Honshu, Japan.
Macroraptorial sperm whales were highly predatory whales of the sperm whale superfamily (Physeteroidea) of the Miocene epoch that hunted large marine mammals, including other whales, using their large teeth. They consist of five genera: Acrophyseter, Albicetus, Brygmophyseter, Livyatan, and Zygophyseter. All species are known by at least a skull, and are informally grouped without a family designation. They were all likely the apex predator of their habitats, comparable to the modern day killer whale, and achieved great lengths, with one species–Livyatan–measuring about 13.5–17.5 m (44–57 ft).
Hoplocetus is an extinct genus of raptorial cetacean of the sperm whale superfamily, Physeteroidea. Its remains have been found in the Miocene of Belgium, France, Germany and Malta, the Pliocene of Belgium and France, and the Pleistocene of the United Kingdom and South Carolina.
Ankylorhiza is an extinct genus of toothed whale that lived in what is now the United States during the Oligocene epoch, between 29 and 23.5 million years ago. The type and only known species is A. tiedemani, though two fossil skeletons may represent an additional, second species within the genus. Ankylorhiza was about 4.8 meters (16 ft) long, with a long, robust skull bearing conical teeth that were angled forwards at the tip of the snout.
Orcas or killer whales have a cosmopolitan distribution and several distinct populations or types have been documented or suggested. Three to five types of orcas may be distinct enough to be considered different races, subspecies, or possibly even species. The IUCN reported in 2008, "The taxonomy of this genus is clearly in need of review, and it is likely that O. orca will be split into a number of different species or at least subspecies over the next few years." Although large variation in the ecological distinctiveness of different orca groups complicate simple differentiation into types. Mammal-eating orcas in different regions were long thought likely to be closely related, but genetic testing has refuted this hypothesis.