A pupa is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages thereof being egg, larva, pupa, and imago. The processes of entering and completing the pupal stage are controlled by the insect's hormones, especially juvenile hormone, prothoracicotropic hormone, and ecdysone. The act of becoming a pupa is called pupation, and the act of emerging from the pupal case is called eclosion or emergence.
The Tachinidae are a large and variable family of true flies within the insect order Diptera, with more than 8,200 known species and many more to be discovered. Over 1,300 species have been described in North America alone. Insects in this family commonly are called tachinid flies or simply tachinids. As far as is known, they all are protelean parasitoids, or occasionally parasites, of arthropods, usually other insects. The family is known from many habitats in all zoogeographical regions and is especially diverse in South America.
The Bombyliidae are a family of flies, commonly known as bee flies. Adults generally feed on nectar and pollen, some being important pollinators. Larvae are mostly parasitoids of other insects.
The apple maggot, also known as the railroad worm, is a species of fruit fly, and a pest of several types of fruits, especially apples. This species evolved about 150 years ago through a sympatric shift from the native host hawthorn to the domesticated apple species Malus domestica in the northeastern United States. This fly is believed to have been accidentally spread to the western United States from the endemic eastern United States region through contaminated apples at multiple points throughout the 20th century. The apple maggot uses Batesian mimicry as a method of defense, with coloration resembling that of the forelegs and pedipalps of a jumping spider.
The Phoridae are a family of small, hump-backed flies resembling fruit flies. Phorid flies can often be identified by their escape habit of running rapidly across a surface rather than taking to the wing. This behaviour is a source of one of their alternate names, scuttle fly. Another vernacular name, coffin fly, refers to Conicera tibialis. About 4,000 species are known in 230 genera. The most well-known species is cosmopolitan Megaselia scalaris. At 0.4 mm in length, the world's smallest fly is the phorid Euryplatea nanaknihali.
The Conopidae, also known as the thick-headed flies, are a family of flies within the Brachycera suborder of Diptera, and the sole member of the superfamily Conopoidea. Flies of the family Conopidae are distributed worldwide in all the biogeographic realms except for the poles and many of the Pacific islands. About 800 species in 47 genera are described worldwide, about 70 of which are found in North America. The majority of conopids are black and yellow, or black and white, and often strikingly resemble wasps, bees, or flies of the family Syrphidae, themselves notable bee mimics. A conopid is most frequently found at flowers, feeding on nectar with its proboscis, which is often long.
Parasitoid wasps are a large group of hymenopteran superfamilies, with all but the wood wasps (Orussoidea) being in the wasp-waisted Apocrita. As parasitoids, they lay their eggs on or in the bodies of other arthropods, sooner or later causing the death of these hosts. Different species specialise in hosts from different insect orders, most often Lepidoptera, though some select beetles, flies, or bugs; the spider wasps (Pompilidae) exclusively attack spiders.
Nasonia are a genus of small pteromalid parasitoid wasps that sting and lay eggs in the pupae of various flies. The fly species that Nasonia usually parasitize are primarily blow flies and flesh flies, making Nasonia a useful tool for biocontrol of these pest flies. The small match-head sized wasps are also referred to as jewel wasps based on the emerald sheen of their exoskeleton.
The fly Megaselia scalaris is a member of the order Diptera and the family Phoridae, and it is widely distributed in warm regions of the world. The family members are commonly known as the "humpbacked fly", the "coffin fly", and the "scuttle fly". The name "scuttle fly" derives from the jerky, short bursts of running, characteristic to the adult fly. The name "coffin fly" is due to their being found in coffins, digging six feet deep in order to reach buried corpses. It is one of the more common species found within the family Phoridae; more than 370 species have been identified within North America.
Bombylius major is a parasitic bee mimic fly. B. major is the most common type of fly within the Bombylius genus. The fly derives its name from its close resemblance to bumblebees and are often mistaken for them.
Trichopoda pennipes is a species of feather-legged fly in the dipteran family Tachinidae.
Compsilura concinnata is a parasitoid native to Europe that was introduced to North America in 1906 to control the population of an exotic forest, univoltine, spongy moth named Lymantria dispar. It is an endoparasitoid of larvae and lives with its host for most of its life. Eventually the parasitoid ends up killing the host and occasionally eating it. It attacks over 200 host species, mainly insects from the Orders: Coleoptera, Lepidoptera and Hymenoptera. Since this parasite has the ability to attack many different types of hosts, the organism has spilled over from the intended forest systems into other areas, like agricultural fields, affecting cabbage pests including the cabbage looper (Trichoplusia); the cabbage worm ; and even other invasive species such as the brown-tail moth. However, it also attacks native, non-pest insects such as the Cecropia moth and American moon moth.
Apocephalus borealis is a species of North American parasitoid phorid fly that attacks bumblebees, honey bees, and paper wasps. This parasitoid's genus Apocephalus is best known for the "decapitating flies" that attack a variety of ant species, though A. borealis attacks and alters the behavior of bees and wasps. These flies are colloquially known as zombie flies and the bees they infect are colloquially known as zombees. Association with honey bees has so far only been documented from California, South Dakota, Oregon, Washington, British Columbia, and Vermont.
Mutilla europaea, the large velvet ant, is a species of parasitoid wasps belonging to the family Mutillidae. It is a parasitoid on various species of bumblebees and is found in Europe, Asia, and North Africa.
Bombus terricola, the yellow-banded bumblebee, is a species of bee in the genus Bombus. It is native to southern Canada and the east and midwest of the United States. It possesses complex behavioral traits, such as the ability to adapt to a queenless nest, choose which flower to visit, and regulate its temperature to fly during cold weather. It was at one time a common species, but has declined in numbers since the late 1990s, likely due to urban development and parasite infection. It is a good pollinator of wild flowers and crops such as alfalfa, potatoes, raspberries, and cranberries.
Macrosiagon limbata is a species of wedge-shaped beetle in the family Ripiphoridae. They are found across the Eastern United States, Southeastern Canada and Central America. It is likely that their range extends into South America but this currently remains unknown.
Gasterophilus intestinalis, also known as horse bot fly, is a species of insect of the family Oestridae, and is found worldwide. The adults, which have a bumblebee-like appearance, are prominently active in the summer. G. intestinalis is primarily a parasite of horses, mules and donkeys.
Liriomyza trifolii, known generally as the American serpentine leafminer or celery leafminer, is a species of leaf miner fly in the family Agromyzidae.
Glossina fuscipes is a riverine fly species in the genus Glossina, which are commonly known as tsetse flies. Typically found in sub-Saharan Africa but with a small Arabian range, G. fuscipes is a regional vector of African trypanosomiasis, commonly known as sleeping sickness, that causes significant rates of morbidity and mortality among humans and livestock. Consequently, the species is among several being targeted by researchers for population control as a method for controlling the disease.
Mallophora ruficauda is a species of parasitic robber fly in the family Asilidae, endemic to South and Central America. Like other robber flies, M. ruficauda is known for its aggressive behavior and predation upon other insects, especially bees. M. ruficauda mimics a bumblebee to fool predators into thinking it has a painful sting and is not worth eating.
