This article duplicates the scope of other articles, specifically Waggle dance. |
This article duplicates the scope of other articles, specifically Tremble dance. |
A round dance is the communicative behaviour of a foraging honey bee ( Apis mellifera ), in which it moves on the comb in close circles, alternating right and then left. [1] It was previously believed that the round dance indicates that the forager has located a profitable food source close to the hive and the round dance transitions into the waggle dance when food sources are more than 50 meters (160 ft) away. [1] Recent research shows that bees have only one dance that always encodes distance and direction to the food source, but that precision and expression of this information depends on the distance to the target; therefore, the use of "round dance" is outdated. [2] Elements of the round dance also provide information regarding the forager's subjective evaluation of the food source's profitability. [3]
Nobel laureate Karl von Frisch was one of the first ethologists to investigate both the waggle dance and round dance through his studies examining honey bee foraging behaviours, and is credited with translating many of their underlying mechanisms. [1]
If a foraging honey bee ( Apis mellifera ) locates a profitable food source, it returns to the hive and performs a round dance to communicate its location. [4] The forager bee moves in close circles over the comb, alternating directions. [1] The round dance is performed by the forager bee when the food source is located in the immediate vicinity of the hive. Karl von Frisch determined that the critical distance for switching between the round dance and the waggle dance exists at 50 meters (160 ft) away from the hive. [1] The scent attached to the forager bee's body communicates the type of food source in question to the follower bees. [1] However, the scent of the food source alone is not sufficient information to guide the follower bees to said food source. [1]
It has been shown that many of the mechanisms used to communicate distance and direction in the waggle dance are also employed in the round dance. [4] The following section will focus on the role of each mechanism as a function of the round dance, specifically.
Acoustic signals are produced as a part of the round dance. These acoustic signals communicate information regarding direction, and give an indication of the distance to the food source. [4] The dance sounds produced by the round dance are airborne, and are of sufficient strength to stimulate Johnston's organs in the antenna of the follower bees to which the forager is communicating. [4] The follower bees extract information about the direction of the food source from the acoustic field that the forager produces. [4] Direction to the food source is communicated through the sound information emitted by the forager bee, as well as through the air currents that accompany the dance sound. [4] Such that, the duration and frequency of the sound produced, along with the magnitude of the pressure gradient produced provide the follower bees with information regarding the direction to the food source. [4] Acoustic signals are also used to communicate distance. [4] The duration of the acoustic signal transmitted is proportional to the distance to the food source. [4] In that, the train of sound pulses produced by the forager bee, known as the dance sound, increases relative to the distance to the food source. [4]
The round dance uses the position of the sun in order to indicate the direction of the food source in the same way that the waggle dance does. [4] On the vertical surface of the comb, the forager expresses the angle between the position of the sun and the path to the food source through an angular deflection from perfect vertical. [1] Honey bees use both the position of the sun and the polarization patterns of a blue sky to communicate the direction to the food source. Support for this theory rests in the observation that honey bees can still recognize the sun's position when it is obscured by a cloud or a mountain, for example. [1] Honey bees also recognize the daily movements of the sun and can use the sun as a compass by calculating the hour of the day. [1]
The round dance communicates the subjective evaluation of a food resource's profitability made by the forager. [3] In regards to nectar, a common food source for honey bees, profitability is determined by its energetic value. [3] Karl von Frisch originally suggested a relationship between what he termed the "liveliness" of the round dance and the value of the food source. [1] Since then, the liveliness of the round dance has been dissected into quantifiable parts. Honey bees communicate information regarding the profitability of a food source through: rate of reversals, number of reversals, and dance duration. [5] Research indicates that the rate of reversals in the round dance is the measure of profitability that is most highly correlated to food source quality. [6] The energetic value of the food source is also correlated with several aspects of the dance sound. There is a positive correlation between energetic value and mean carrier frequency, pulse repetition rate, amplitude, and duration of the sound bursts. [5]
However, costs can be associated with the food sources, and consequently lower their subjective profitability. Some possible costs include: distance from the hive, and nectar flow rate. [3] Furthermore, the forager's subjective rating of the profitability of a food source depends on the quality of food sources that the forager had previously encountered. If the forager had previously encountered a food source of higher energetic value, and less cost, it will rate the current food source as less profitable than if it had previously encountered a food source of less energetic value. [3] Thus, the forager bee integrates the costs and benefits associated with each potential food source and communicates their subjective profitability through their round dance performance. However, the forager bee only communicates the food source if it passes a certain concentration threshold for dancing, by passing this threshold it is determined that there are enough benefits associated with the food source to justify expending the energy required for performing the round dance. [6]
Although the round dance communicates the different profitabilities associated with each food source, the round dances displaying both rich and poor food resources are equally stimulating to the follower bees. [7] The follower bees do not respond to only the most rich food resources because the benefits of being highly selective about which dance to respond to are low. [7] It is suggested that honey bees benefit more from responding to a wide array of food sources rather than all congregating at the same, slightly richer food source. [7] Also, since all food resources have to pass the concentration threshold for dancing in order to be communicated via the round dance, any food resource that is communicated through the round dance must have a relatively high profitability. [6] [7]
A honey bee is a eusocial flying insect within the genus Apis of the bee clade, all native to mainland Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the current cosmopolitan distribution of honey bees, introducing multiple subspecies into South America, North America, and Australia.
Karl Ritter von Frisch, was a German-Austrian ethologist who received the Nobel Prize in Physiology or Medicine in 1973, along with Nikolaas Tinbergen and Konrad Lorenz.
Bee learning and communication includes cognitive and sensory processes in all kinds of bees, that is the insects in the seven families making up the clade Anthophila. Some species have been studied more extensively than others, in particular Apis mellifera, or European honey bee. Color learning has also been studied in bumblebees.
Apis mellifera ligustica is the Italian bee or the Italian Honey bee which is a subspecies of the western honey bee.
For bees, their forage or food supply consists of nectar and pollen from blooming plants within their flight range. The forage sources for honey bees are an important consideration for beekeepers. In order to determine where to locate hives for maximum honey production and brood one must consider the off-season. If there are no honey flows the bees may have to be fed. Bees that are used for commercial pollination are usually fed in the holding yards. Forage is also significant for pollination management with other bee species. Nectar contains sugars that are the primary source of energy for the bees' wing muscles and for heat for honey bee colonies during the winter. Pollen provides the protein and trace minerals that are mostly fed to the brood in order to replace bees lost in the normal course of their life cycle and colony activity.
Waggle dance is a term used in beekeeping and ethology for a particular figure-eight dance of the honey bee. By performing this dance, successful foragers can share information about the direction and distance to patches of flowers yielding nectar and pollen, to water sources, or to new nest-site locations with other members of the colony.
The dwarf honey bee, Apis florea, is one of two species of small, wild honey bees of southern and southeastern Asia. It has a much wider distribution than its sister species, Apis andreniformis. First identified in the late 18th century, Apis florea is unique for its morphology, foraging behavior and defensive mechanisms like making a piping noise. Apis florea have open nests and small colonies, which makes them more susceptible to predation than cavity nesters with large numbers of defensive workers. These honey bees are important pollinators and therefore commodified in countries like Cambodia.
The Cape honey bee or Cape bee is a southern South African subspecies of the western honey bee. They play a major role in South African agriculture and the economy of the Western Cape by pollinating crops and producing honey in the Western Cape region of South Africa. The species is endemic to the Western Cape region of South Africa on the coastal side of the Cape Fold mountain range.
Apis andreniformis, or the black dwarf honey bee, is a relatively rare species of honey bee whose native habitat is the tropical and subtropical regions of Southeast Asia.
Apis dorsata, the rock bee or giant honey bee, is a honey bee of South and Southeast Asia. They are typically around 17–20 mm (0.7–0.8 in) long and nests are mainly built in exposed places far off the ground, like on tree limbs, under cliff overhangs, and under buildings. These social bees are known for their aggressive defense strategies and vicious behavior when disturbed. Though not domesticated, indigenous peoples have traditionally used this species as a source of honey and beeswax, a practice known as honey hunting.
A tremble dance is a dance performed by forager honey bees of the species Apis mellifera to recruit more receiver honey bees to collect nectar from the workers.
The East African lowland honey bee is a subspecies of the western honey bee. It is native to central, southern and eastern Africa, though at the southern extreme it is replaced by the Cape honey bee. This subspecies has been determined to constitute one part of the ancestry of the Africanized bees spreading through North and South America.
In computer science and operations research, the bees algorithm is a population-based search algorithm which was developed by Pham, Ghanbarzadeh et al. in 2005. It mimics the food foraging behaviour of honey bee colonies. In its basic version the algorithm performs a kind of neighbourhood search combined with global search, and can be used for both combinatorial optimization and continuous optimization. The only condition for the application of the bees algorithm is that some measure of distance between the solutions is defined. The effectiveness and specific abilities of the bees algorithm have been proven in a number of studies.
The western honey bee or European honey bee is the most common of the 7–12 species of honey bees worldwide. The genus name Apis is Latin for 'bee', and mellifera is the Latin for 'honey-bearing' or 'honey-carrying', referring to the species' production of honey.
Apis cerana japonica is a subspecies of the eastern honeybee native to Japan. It is commonly known as the Japanese honeybee. Analysis of mitochondrial DNA suggests that the ancestors of this subspecies came to Japan from the Korean Peninsula via Tsushima Island. Genetic differentiation between Japanese honeybees and Korean honeybees occurred about 20,000 years ago, which coincides with the separation of Japan's Tsushima Island from the Korean Peninsula due to sea level rise. They have been observed moving into urban areas in the absence of natural predators.
Apis cerana, the eastern honey bee, Asiatic honey bee or Asian honey bee, is a species of honey bee native to South, Southeast and East Asia. This species is the sister species of Apis koschevnikovi and both are in the same subgenus as the western (European) honey bee, Apis mellifera. A. cerana is known to live sympatrically along with Apis koschevnikovi within the same geographic location. Apis cerana colonies are known for building nests consisting of multiple combs in cavities containing a small entrance, presumably for defense against invasion by individuals of another nest. The diet of this honey bee species consists mostly of pollen and nectar, or honey. Moreover, Apis cerana is known for its highly social behavior, reflective of its classification as a type of honey bee.
In linguistics, displacement is the capability of language to communicate about things that are not immediately present ; i.e., things that are either not here or are not here now.
Bumblebees, like the honeybee collect nectar and pollen from flowers and store them for food. Many individuals must be recruited to forage for food to provide for the hive. Some bee species have highly developed ways of communicating with each other about the location and quality of food resources ranging from physical to chemical displays.
Sniffer bees or sniffer wasps are insects in the order Hymenoptera that can be trained to perform a variety of tasks to detect substances such as explosive materials or illegal drugs, as well as some human and plant diseases. The sensitivity of the olfactory senses of bees and wasps in particular have been shown to rival the abilities of sniffer dogs, though they can only be trained to detect a single scent each.
Hockett's Design Features are a set of features that characterize human language and set it apart from animal communication. They were defined by linguist Charles F. Hockett in the 1960s. He called these characteristics the design features of language. Hockett originally believed there to be 13 design features. While primate communication utilizes the first 9 features, the final 4 features are reserved for humans. Hockett later added prevarication, reflexiveness, and learnability to the list as uniquely human characteristics. He asserted that even the most basic human languages possess these 16 features.
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