The Ladinian is a stage and age in the Middle Triassic series or epoch. It spans the time between 242 Ma and ~237 Ma. The Ladinian was preceded by the Anisian and succeeded by the Carnian.
Cymbospondylus is an extinct genus of large ichthyosaurs, of which it is among the oldest representatives, that lived from the Lower to Middle Triassic in what are now North America and Europe. The first known fossils of this taxon are a set of more or less complete vertebrae which were discovered in the 19th century in various mountain ranges of Nevada, in the United States, before being named and described by Joseph Leidy in 1868. It is in the beginning of the 20th century that more complete fossils were discovered through several expeditions launched by the University of California, and described in more detail by John Campbell Merriam in 1908, thus visualizing the overall anatomy of the animal. While many species have been assigned to the genus, only five are recognized as valid, the others being considered synonymous, doubtful or belonging to other genera. Cymbospondylus was formerly classified as a representative of the Shastasauridae, but more recent studies consider it to be more basal, view as the type genus of the Cymbospondylidae.
Ticinosuchus is an extinct genus of suchian archosaur from the Middle Triassic of Switzerland and Italy.
Monte San Giorgio is a Swiss mountain and UNESCO World Heritage Site near the border between Switzerland and Italy. It is part of the Lugano Prealps, overlooking Lake Lugano in the Swiss Canton of Ticino.
Ticinepomis is an extinct genus of coelacanth lobe-finned fish which lived during the Middle Triassic period in what is now Switzerland. It contains two species, T. peyeri and T. ducanensis.
Besanosaurus is a extinct genus of Middle Triassic ichthyosaur from Monte San Giorgio of Italy and Switzerland, containing the single species B. leptorhynchus. Besanosaurus was named by Cristiano Dal Sasso and Giovanni Pinna in 1996, based on the nearly complete flattened skeleton BES SC 999, the holotype specimen. This skeleton is preserved across multiple thin rock slabs spanning 3.5 by 4 metres when assembled and took thousands of hours to prepare. Additional specimens from Monte San Giorgio that have previously been considered separate genera, including a partial skull named Mikadocephalus and a well-preserved, largely complete skeleton, have been reinterpreted as additional specimens of Besanosaurus. Putative specimens of Besanosaurus have been discovered in the Norwegian archipelago of Svalbard and Germany, although their attribution to this genus remains disputed.
Serpianosaurus is an extinct genus of pachypleurosaurs known from the Middle Triassic deposits of Switzerland and Germany. It was a small reptile, with the type specimen of S. mirigiolensis measuring 75 cm (2.46 ft) long.
Luganoia is an extinct genus of prehistoric bony fish that lived during the Anisian and Ladinian ages of the Middle Triassic epoch. Fossils were recovered from the Besano Formation of Monte San Giorgio and Besano area and from the Zhuganpo Member of Guizhou, South China. It was also reported from the Ladinian of Spain.
Besania is an extinct genus of prehistoric marine ray-finned fish that lived during the Anisian and Ladinian ages of the Middle Triassic epoch in what is now southern/southeastern Switzerland and northern Italy. Fossils were recovered from the Besano Formation of Monte San Giorgio area and the Prosanto Formation of canton Graubünden, Switzerland.
Ctenognathichthys is an extinct genus of prehistoric marine ray-finned fish that lived during the Middle Triassic epoch of Europe, in the former Tethys Ocean.
Heptanema is an extinct genus of prehistoric coelacanth from the Middle Triassic (Ladinian) of northern Italy and southern Switzerland.
Acentrophorus is an extinct genus of prehistoric freshwater and marine ray-finned fish from the Roadian to the Wuchiapingian of England, Germany (Kupferschiefer), Italy and Russia. There may also be a Triassic occurrence in Australia.
Colobodus is an extinct genus of marine Triassic ray-finned fish of the family Colobodontidae and order Perleidiformes. Fossils have been found in Europe and China, encompassing the former Tethys Ocean. It could reach body lengths of about 70 cm.
Meridensia is an extinct genus of prehistoric ray-finned fish that lived during the Anisian and Ladinian ages of the Middle Triassic epoch in what is now southern Switzerland and northern Italy. Fossils were recovered from the Besano Formation of Monte San Giorgio and Besano area at the Swiss-Italian boundary.
Wimanius is a genus of ichthyosaur from the Middle Triassic of Switzerland, containing a single species, Wimanius odontopalatus. It was described by Michael Maisch and Andreas Matzke in 1998 based on an incomplete skull from Monte San Giorgio, a mountain on the Swiss-Italian border. Wimanius possesses teeth on its palate, though whether they were located on the palatine or pterygoid is disputed. Other features of Wimanius include a large orbit and jugals with two rami of similar lengths. Different phylogenetic placements of Wimanius have been recovered by different studies, including it being a mixosaurid relative or a merriamosaur, and a monotypic family, Wimaniidae has been named for it. However, its validity has also been questioned, and synonymy with various other genera has been proposed. The only specimen of Wimanius come from the Besano Formation. During the Anisian, this region was a lagoon populated by a wide variety of marine life, including a variety of other ichthyosaurs.
The Wetterstein Formation is a regional geologic formation of the Northern Limestone Alps and Western Carpathians extending from southern Bavaria, Germany in the west, through northern Austria to northern Hungary and western Slovakia in the east. The formation dates back to the Ladinian to Carnian stages of the Late Triassic. The formation is named after the Wetterstein Mountains in southern Germany and northwestern Austria. The center of its distribution, however, is in the Karwendel Mountains. It occurs in the Northern and Southern Limestone Alps and in the Western Carpathians.
Eusaurosphargis is an extinct genus of a diapsid reptile, known from the Middle Triassic Besano Formation of northern Italy and Prosanto Formation of south-eastern Switzerland. It contains a single species, Eusaurosphargis dalsassoi. It was a small reptile, measuring 20 cm (7.9 in) long.
The Besano Formation is a geological formation in the southern Alps of northwestern Italy and southern Switzerland. This formation, a thin but fossiliferous succession of dolomite and black shale, is famous for its preservation of Middle Triassic (Anisian–Ladinian) marine life including fish and aquatic reptiles. It is exposed in the Monte San Giorgio and Besano area. It is among the formations responsible for the area being designated as a UNESCO World Heritage Site. In Switzerland, it is also known as the Grenzbitumenzone. The Anisian-Ladinian boundary lies in the upper part of the Besano Formation.
Acronemus is an extinct genus of euselachian from the Middle Triassic of Switzerland. It is an enigmatic genus with uncertain relations to other groups. Though originally placed within Ctenacanthiformes, it is now considered Euselachii incertae sedis, due to its mixture of features similar to hybodontiforms and neoselachians. Originally, teeth from this genus were attributed to "Acrodus bicarinatus" while fin spines were named "Nemacanthus tuberculatus". Associated material showed they were the same animal, with the older specific epithet (tuberculatus) taking precedence. The euselachian was given the new genus Acrocnemus, containing a single species. Acronemus is found in the Anisian-age Grenzbitumenzone of Monte San Giorgio. It was a small animal measuring 30–35 cm (0.98–1.15 ft) long. A 2018 study considered it to be closely related to the early Carboniferous genus Tristychius as part of the family Tristychiidae as a basal euselachian, with hybodonts more closely related to neoselachians than to Tristychiidae.
Marcopoloichthys is an extinct genus of marine teleosteomorph ray-finned fish known from the Middle and Late Triassic of the former Tethys Ocean. It is the only genus in the family Marcopoloichthyidae. It was originally described based on specimens from both Italy and China, hence the name Marcopoloichthys, which references the medieval Venetian merchant Marco Polo, who traveled from Italy to China.
Italy 
Switzerland 
