Ulmus 'Columella' | |
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![]() 'Columella', Amsterdam. | |
Genus | Ulmus |
Hybrid parentage | 'Plantyn' selfed or openly pollinated |
Cultivar | 'Columella' |
Origin | Wageningen, The Netherlands |
Ulmus 'Columella' is a Dutch elm cultivar raised by the Dorschkamp Research Institute for Forestry & Landscape Planning, Wageningen. Originally thought to have been derived from a selfed or openly pollinated seedling of the hybrid clone 'Plantyn', DNA analysis later determined it arose from a cross of 'Plantyn' and Ulmus minor . Sown in 1967, it was released for sale in 1989 after proving extremely resistant to Dutch elm disease following injection with unnaturally high doses of the pathogen, Ophiostoma novo-ulmi . [1] [2] However, propagated by grafting onto wych elm rootstocks, graft failure owing to incompatibility has become a common occurrence in the Netherlands.
'Columella' makes a tall, fastigiate tree with very upright branches, but broadens in later years . The rough, rounded, and twisted leaves, < 7 cm (3 in) long, are the result of a recessive gene inherited from its Exeter elm ancestor, and are arranged in asymmetric clusters on short branchlets. The samarae, broadly obovate, are 13–17 mm (0.5–0.7 in) long by 10–12 mm wide.
Rated 5 out of 5, [3] 'Columella' has a very high resistance to Dutch elm disease.
Wind resistant, the tree has been planted throughout the Netherlands, where its columnar shape has made it popular as a street tree. It is commonly found in Amsterdam, where it has been widely planted as a replacement for the similarly fastigiate Guernsey Elm, U. minor 'Sarniensis' , itself a replacement for the Belgian Elm, Ulmus × hollandica 'Belgica', which had succumbed so readily to the earlier strain of Dutch elm disease after World War I. 'Columella' has also been planted to replace Guernsey Elm in Edinburgh. [4]
In trials conducted by Butterfly Conservation in southern Hampshire, England, 'Columella' was the only cultivar to become distressed during the drought of 2006, shedding most of its foliage by early August; a trait possibly inherited from one of the tree's ancestors, the Himalayan elm Ulmus wallichiana . 'Columella' first flowered aged 8 years, in March; the resultant seeds were found to have a moderate viability. [5] [6] 'Columella' featured in New Zealand government trials during the 1990s at the Hortresearch station, Palmerston North.
The hybrid is named for the Roman agronomist Columella, who introduced the Atinian elm (now more commonly known as the English elm) to Spain from Italy circa c. AD 50.
Ulmus 'San Zanobi' is a hybrid elm cultivar raised by the Istituto per la Protezione delle Piante (IPP) in Florence, from a crossing of the Dutch hybrid 'Plantyn' and the Siberian Elm Ulmus pumila clone 'S 15'; it was released to commerce in 2002. 'San Zanobi' was introduced to the UK in 2004 by Hampshire & Isle of Wight Branch, Butterfly Conservation, as part of its assessment of DED-resistant cultivars as potential hosts of the endangered White-letter Hairstreak.
Ulmus 'Plinio' is a hybrid elm cultivar derived from a crossing of the Dutch cultivar 'Plantyn' with the Siberian Elm Ulmus pumila clone 'S.2'. 'Plinio' was raised by the Istituto per la Protezione delle Piante (IPP), in Florence and released for sale in 2003. 'Plinio' was introduced to the UK in 2004 by Hampshire & Isle of Wight Branch, Butterfly Conservation, as part of its assessment of DED-resistant cultivars as potential hosts of the endangered White-letter Hairstreak.
Ulmus davidiana var. japonica, the Japanese elm, is one of the larger and more graceful Asiatic elms, endemic to much of continental northeast Asia and Japan, where it grows in swamp forest on young alluvial soils, although much of this habitat has now been lost to intensive rice cultivation.
Ulmus 'Nanguen' is a complex fourth generation Dutch hybrid cultivar raised at the Dorschkamp Research Institute for Forestry & Landscape Planning, Wageningen. Lutèce was derived from the cross 'Plantyn' ×, an ancestry comprising four field elms, a wych elm, the curious Exeter Elm ('Exoniensis'), and a frost-resistant selection of the Himalayan elm.
Ulmus 'New Horizon' is an American hybrid cultivar raised by the Wisconsin Alumni Research Foundation (WARF), from a crossing of the Japanese Elm clone W43-8 = 'Reseda' with Siberian Elm clone W426 grown from seed collected from a street tree at Yankton, South Dakota. As selection 'W917' it is a sibling of 'Rebona'. Tested in the US National Elm Trial coordinated by Colorado State University, 'New Horizon' averaged a survival rate of 74% after 10 years. 'New Horizon' was patented in the US in 1994, while in Europe, it is marketed as one of the 'Resista' elms protected under EU breeders' rights.
Ulmus × hollandica 'Commelin' is a Dutch hybrid cultivar released for sale in 1960. The tree was raised at Baarn as clone 274 by the Foundation Willie Commelin Scholten Phytopathological Laboratory in 1940, from a crossing of Ulmus × hollandica 'Vegeta' and clone 1, an Ulmus minor selected from a 1929 elm seedlings lot obtained from the Barbier nursery, Orléans.
The Dutch hybrid elm cultivar Ulmus × hollandica 'Groeneveld' was cloned in 1949 at the De Dorschkamp Institute, Wageningen, and released in 1963 in response to the earlier, less virulent form of Dutch elm disease that afflicted Europe shortly after the First World War. The cultivar was derived from a crossing of Dutch clones '49', and '1', a Field Elm Ulmus minor found in central France and marketed by the Barbier nursery in Orléans.
Ulmus 'Lobel' is a Dutch hybrid cultivar raised at the Dorschkamp Research Institute for Forestry & Landscape Planning, Wageningen, from a crossing of clone '202' with '336'. 'Lobel' was cloned in 1962 and released for sale in 1973.
The hybrid elm cultivar Ulmus × hollandica 'Pioneer' is an American clone arising from the crossing of two European species, Wych Elm U. glabra and Field Elm U. minor. Raised by the USDA station at Delaware, Ohio, in 1971, 'Pioneer' was released to commerce in 1983.
Ulmus 'Plantyn' was one of three Dutch hybrid elms released by the Dorschkamp Research Institute for Forestry & Landscape Planning, Wageningen, in 1973. Derived from a crossing of the Dutch hybrids '202' and '302', it was to prove of great significance in later developments. A selfed seedling was to become the first Dutch clone to prove effectively immune to disease, released in 1989 as 'Columella'. 'Plantyn' was also destined to be the female parent of Lutèce released in 2002. In Italy, 'Plantyn' was used again as female parent in hybridizations with the Siberian Elm Ulmus pumila by the Istituto per la Protezione delle Piante (IPP), to create three new cultivars better adapted to the Mediterranean climate.
Ulmus 'Wanoux' is a Dutch hybrid cultivar raised at the Dorschkamp Research Institute for Forestry & Landscape Planning, Wageningen, from an open pollination of 'Plantyn'. Originally identified as clone No. 762, it was selected for assessment by the French Institut National de la Recherche Agronomique (INRA), which patented it as 'Wanoux' in 2006.
The Japanese elm cultivar Ulmus davidianavar.japonica 'Prospector' was originally treated as a cultivar of Wilson's elm U. wilsonianaSchneid., a species sunk as Ulmus davidiana var. japonica by Fu. A U.S. National Arboretum introduction, it was selected in 1975 from a batch of 1965 seedlings in Delaware, Ohio, and released without patent restrictions in 1990. 'Prospector' proved moderately successful in the US National Elm Trial, averaging a survival rate of 76% overall.
Ulmus 'Patriot' is a hybrid cultivar raised by the United States National Arboretum in 1980. Derived from a crossing of the American hybrid 'Urban' with the Wilson's Elm cultivar 'Prospector', 'Patriot' was released to commerce, free of patent restrictions, in 1993. Tested in the US National Elm Trial coordinated by Colorado State University, 'Patriot' averaged a survival rate of 85% after 10 years.
Ulmus 'Arno' is an Italian hybrid cultivar derived from a crossing of 'Plantyn' with the Siberian Elm Ulmus pumila clone S.2. It was raised by the Istituto per la Protezione delle Piante (IPP) in Florence, and released in 2007. However, 'Arno' was not a commercial success; propagation had ceased by 2010, and it is no longer patent protected.
Ulmus 'Fiorente' is a hybrid cultivar elm derived from a crossing of the Siberian Elm Ulmus pumila clone 'S.10' from Lucca, Italy, with the Ulmus minor clone 'C.02' from Lungarno, Florence, by the Istituto per la Protezione delle Piante (IPP), part of the Italian National Research Council, in Florence. The tree is protected by Plant Breeders' Rights bestowed by the EU on 25 March 2010.
Ulmus laciniata var. nikkoensisRehder, the Nikko elm, was discovered as a seedling near Lake Chūzenji, near Nikkō, Japan, and obtained by the Arnold Arboretum in 1905. The taxonomy of the tree remains a matter of contention, and has been considered possibly a hybrid of U. laciniata and U. davidiana var. japonica. However, in crossability experiments at the Arnold Arboretum in the 1970s, U. laciniata, a protogynous species, was found to be incompatible with U. davidiana var. japonica, which is protandrous.
Ulmus 'Rebona' is an American hybrid cultivar raised by the Wisconsin Alumni Research Foundation (WARF) as selection 'W916', derived from a crossing of Japanese Elm clone W43-8 = 'Reseda' with Siberian Elm clone W426 grown from seed collected from a street tree at Yankton, South Dakota. The tree was registered in 1993 by Conrad Appel KG, of Darmstadt and is a sibling of 'New Horizon'. In Europe, 'Rebona' is marketed as a Resista elm protected under E U breeders' rights.
Ulmus 'Morfeo' is a hybrid elm cultivar raised by the Istituto per la Protezione delle Piante (IPP), Florence, in 2000. 'Morfeo' arose from a crossing of the Dutch hybrid clone '405' and the Chenmou Elm, the latter a small tree from the provinces of Anhui and Jiangsu in eastern China, The '405' clone is a full sister of 'Groeneveld', a crossing of an English U. × hollandica and a French U. minor from the Barbier Nursery, Orléans.
Ulmus 'Wingham' is a complex hybrid elm cultivar featuring two Asiatic and two European species, bestowing it with an exceptionally high resistance to Dutch elm disease (DED). It was raised as clone no. FL493 by the Istituto per la Protezione delle Piante (IPP) in Florence, but never patented owing to its limited aesthetic appeal. It was introduced to the UK in 2011 by David Herling, Resistant Elms, who trialled it successfully at Wingham in Kent. It was later trialled by Hampshire & Isle of Wight Branch, Butterfly Conservation, as part of an assessment of DED-resistant cultivars as potential hosts of the endangered White-letter Hairstreak. 'Wingham' was released for sale in the UK in 2019.