Umkomasia feistmantelii Temporal range: | |
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Umkomasia feistmantelii ovulate structure from the Early Triassic Newport Formation of Turimetta Head, NSW, Australia. | |
Scientific classification ![]() | |
Kingdom: | Plantae |
Clade: | Tracheophytes |
Order: | † Peltaspermales |
Family: | † Corystospermaceae |
Genus: | † Umkomasia |
Species: | †U. feistmantelii |
Binomial name | |
†Umkomasia feistmantelii Holmes 1987 | |
Umkomasia feistmantelii is an unusually large species of Umkomasia from the Early Triassic of New South Wales, Australia.
Umkomasia feistmantelii is found both with cupules enclosing the large seeds and with cupules open and expandede into a star-shaped form.
Umkomasia feistmantelii from the Early Triassic of Australia may have been produced by the same plant as Pteruchus barrealensis (pollen organs) and Dicroidium zuberi (leaves) [1]
The Mesozoic Era, also called the Age of Reptiles and the Age of Conifers, is the second-to-last era of Earth's geological history, lasting from about 252 to 66 million years ago and comprising the Triassic, Jurassic and Cretaceous Periods. It is characterized by the dominance of archosaurian reptiles, like the dinosaurs; an abundance of conifers and ferns; a hot greenhouse climate; and the tectonic break-up of Pangaea. The Mesozoic is the middle of the three eras since complex life evolved: the Paleozoic, the Mesozoic, and the Cenozoic.
The Phanerozoic Eon is the current geologic eon in the geologic time scale, and the one during which abundant animal and plant life has existed. It covers 538.8 million years to the present, and it began with the Cambrian Period, when animals first developed hard shells preserved in the fossil record. The time before the Phanerozoic, called the Precambrian, is now divided into the Hadean, Archaean and Proterozoic eons.
The Permian–Triassicextinction event, also known as the End-Permian Extinction and colloquially as the Great Dying, formed the boundary between the Permian and Triassic geologic periods, as well as between the Paleozoic and Mesozoic eras, approximately 251.9 million years ago. It is the Earth's most severe known extinction event, with the extinction of 57% of biological families, 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species. It was the largest known mass extinction of insects.
The Triassic is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago (Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic.
The Triassic–Jurassic (Tr-J) extinction event, often called the end-Triassic extinction, marks the boundary between the Triassic and Jurassic periods, 201.3 million years ago, and is one of the major extinction events of the Phanerozoic eon, profoundly affecting life on land and in the oceans. In the seas, the entire class of conodonts and 23–34% of marine genera disappeared. On land, all archosauromorphs other than crocodylomorphs, pterosaurs, and dinosaurs became extinct; some of the groups which died out were previously abundant, such as aetosaurs, phytosaurs, and rauisuchids. Some remaining non-mammalian therapsids and many of the large temnospondyl amphibians had become extinct prior to the Jurassic as well. However, there is still much uncertainty regarding a connection between the Tr-J boundary and terrestrial vertebrates, due to a lack of terrestrial fossils from the Rhaetian (latest) stage of the Triassic. What was left fairly untouched were plants, dinosaurs, pterosaurs and mammals; this allowed the dinosaurs and pterosaurs to become the dominant land animals for the next 135 million years.
Glossopteris [etymology: from Ancient Greek γλῶσσα + πτερίς ] is the largest and best-known genus of the extinct Permian order of seed ferns known as Glossopteridales. The genus Glossopteris refers only to leaves, within a framework of form genera used in paleobotany. Species of Glossopteris were the dominant trees of the middle- to high-latitude lowland vegetation across the supercontinent Gondwana during the Permian Period. Glossopteris fossils were critical in recognizing former connections between the various fragments of Gondwana: South America, Africa, India, Australia, New Zealand, and Antarctica.
In the geologic timescale, the Middle Triassic is the second of three epochs of the Triassic period or the middle of three series in which the Triassic system is divided in chronostratigraphy. The Middle Triassic spans the time between 247.2 Ma and 237 Ma. It is preceded by the Early Triassic Epoch and followed by the Late Triassic Epoch. The Middle Triassic is divided into the Anisian and Ladinian ages or stages.
Gondwana was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic and began to break up during the Jurassic period. The final stages of break-up, involving the separation of Antarctica from South America and Australia, occurred during the Paleogene. Gondwana was not considered a supercontinent by the earliest definition, since the landmasses of Baltica, Laurentia, and Siberia were separated from it. To differentiate it from the Indian region of the same name, it is also commonly called Gondwanaland.
Brachyopoidea is a superfamily of temnospondyls that lived during the Mesozoic. It contains the families Brachyopidae and Chigutisauridae. The earliest records of brachyopids are from the Lower Triassic in Australia. The latest-surviving member of the superfamily is the chigutisaurid Koolasuchus from the Early Cretaceous of Australia.
The Caytoniales are an extinct order of seed plants known from fossils collected throughout the Mesozoic Era, around 252 to 66 million years ago. They are regarded as seed ferns because they are seed-bearing plants with fern-like leaves. Although at one time considered angiosperms because of their berry-like cupules, that hypothesis was later disproven. Nevertheless, some authorities consider them likely ancestors or close relatives of angiosperms. The origin of angiosperms remains unclear, and they cannot be linked with any known seed plants groups with certainty.
The Lyginopteridales were the archetypal pteridosperms: They were the first plant fossils to be described as pteridosperms and, thus, the group on which the concept of pteridosperms was first developed; they are the stratigraphically oldest-known pteridosperms, occurring first in late Devonian strata; and they have the most primitive features, most notably in the structure of their ovules. They probably evolved from a group of Late Devonian progymnosperms known as the Aneurophytales, which had large, compound frond-like leaves. The Lyginopteridales became the most abundant group of pteridosperms during Mississippian times, and included both trees and smaller plants. During early and most of middle Pennsylvanian times the Medullosales took over as the more important of the larger pteridosperms but the Lyginopteridales continued to flourish as climbing (lianescent) and scrambling plants. However, later in Middle Pennsylvanian times the Lyginopteridales went into serious decline, probably being out-competed by the Callistophytales that occupied similar ecological niches but had more sophisticated reproductive strategies. A few species continued into Late Pennsylvanian times, and in Cathaysia and east equatorial Gondwana they persisted into the Late Permian, but subsequently became extinct. Most evidence of the Lyginopteridales suggests that they grew in tropical latitudes of the time, in North America, Europe and China.
Dicroidium is an extinct genus of fork-leaved seed ferns that were widely distributed over Gondwana during the Triassic. Their fossils are known from South Africa, the Arabian Peninsula, Australia, New Zealand, South America, Madagascar, the Indian subcontinent and Antarctica. They were first discovered in Triassic sediments of Tasmania by Morris in 1845. Fossils from the Umm Irna Formation in Jordan and in Pakistan indicate that these plants already existed in Late Permian. Late surviving members of the genus are known from the Early Jurassic (Sinemurian) of East Antarctica. Within paleobotany, Dicroidium is a form genus used to refers to the leaves, associated with ovuluate organs classified as Umkomasia and pollen organs classified as Pteruchus, while Dicroidum is also used collectively to refer to the whole plant.
Umkomasia is a genus of seed bearing organs produced by corystosperm seed ferns, first based on fossils collected by Hamshaw Thomas from the Burnera Waterfall locality near the Umkomaas River of South Africa. He recognized on the basis of cuticular similarities that the same plant produced pollen organs Pteruchus and the leaves Dicroidium. Various other corystosperm seed bearing organs from the Jurassic and Cretaceous have been assigned to this genus, but recently have been given distinct genera, with Umkomasia being restricted to the Triassic.
Dicroidium odontopteroides was a common and widespread species of Dicroidium known from South Africa, Australia, New Zealand, South America and Antarctica. The species was first discovered in Triassic sediments of Tasmania and described by the palaeontologist John Morris in 1845.
Pteruchus barrealensis is an unusually large species of Pteruchus with very elongate polleniferous heads from Early Triassic of Australia and Argentina.
Dicroidium zuberi is a large bipinnate species of the seed fern Dicroidium with a forked rachis. The leaves are affiliated with Umkomasia feistmantellii megasporophylls and Petruchusbarrealensis microsporophylls.
Pteruchus is a form genus for pollen organs of the seed fern (Pteridospermatophyta family Umkomasiaceae. It was first described by Hamshaw Thomas from the Umkomaas locality of South Africa. It is associated with the seed bearing organs Umkomasia and Dicroidium leaves.
Peltaspermaceae is a natural family of seed ferns (Pteridospermatophyta) widespread in both northern and southern hemispheres coal measures of Permian and Triassic age. Peltasperms persisted in a relictual distribution in Patagonia during the Early Jurassic.
Corystospermaceae is a natural family of seed ferns (Pteridospermatophyta) also called Umkomasiaceae, and first based on fossils collected by Hamshaw Thomas from the Burnera Waterfall locality near the Umkomaas River of South Africa The leaves of Dicroidium were recognized by Alex Du Toit to unite all the countries of the Gondwana supercontinent during the Triassic: Africa, South America, India, and Australia. Subsequently, Dicroidium was found in the Triassic of Antarctica and New Zealand, and also the Permian Umm Irna Formation of Jordan. According to the form generic system of paleobotany, leaves are given separate generic names to ovulate and pollen organs, but the discovery of these reproductive organs in Africa by Thomas, and subsequently throughout Gondwana, strengthened Du Toit's concept of a continuous southern supercontinent. Corystospermaceae were also components of Jurassic and Cretaceous floras, declining in the Cretaceous presumably due to the rise of flowering plants, the last known representative of the group, Komlopteris cenozoicus, is known from the Eocene of Tasmania, at least 13 million years after the Cretaceous–Paleogene extinction event.
Rock cupules are artificially made depressions on rock surfaces that resemble the shape of an inverse spherical cap or dome. They were made by direct percussion with hand-held hammer-stones, on vertical, sloping or horizontal rock surfaces. Cupules are widely believed to be the world's most common rock art motifs, found in huge numbers in every continent except Antarctica. They were produced in many cultures, from the Lower Paleolithic to the 20th century, and they can be found on most lithologies. Similar artifacts from lithic Native American cultures are also known as cupstones.