You can help expand this article with text translated from the corresponding article in German. (January 2024)Click [show] for important translation instructions.
|
Felidae [1] | |
---|---|
Clockwise, a tiger (Panthera tigris), Canada lynx (Lynx canadensis), fishing cat (Prionailurus viverrinus), European wildcat (Felis silvestris), ocelot (Leopardus pardalis), Asian golden cat (Catopuma temminckii), serval (Leptailurus serval), and cougar (Puma concolor). | |
Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Chordata |
Class: | Mammalia |
Order: | Carnivora |
Suborder: | Feliformia |
Superfamily: | Feloidea |
Family: | Felidae Fischer von Waldheim, 1817 |
Type genus | |
Felis | |
Genera and Subfamilies | |
| |
The native distribution and density of extant felid species. |
Felidae ( /ˈfɛlɪdiː/ ) is the family of mammals in the order Carnivora colloquially referred to as cats. A member of this family is also called a felid ( /ˈfiːlɪd/ ). [3] [4] [5] [6] The term "cat" refers both to felids in general and specifically to the domestic cat ( Felis catus). [7]
The 41 extant Felidae species exhibit the greatest diversity in fur patterns of all terrestrial carnivores. [8] Cats have retractile claws, slender muscular bodies and strong flexible forelimbs. Their teeth and facial muscles allow for a powerful bite. They are all obligate carnivores, and most are solitary predators ambushing or stalking their prey. Wild cats occur in Africa, Europe, Asia and the Americas. Some wild cat species are adapted to forest and savanna habitats, some to arid environments, and a few also to wetlands and mountainous terrain. Their activity patterns range from nocturnal and crepuscular to diurnal, depending on their preferred prey species. [9]
Reginald Innes Pocock divided the extant Felidae into three subfamilies: the Pantherinae, the Felinae and the Acinonychinae, differing from each other by the ossification of the hyoid apparatus and by the cutaneous sheaths which protect their claws. [10] This concept has been revised following developments in molecular biology and techniques for the analysis of morphological data. Today, the living Felidae are divided into two subfamilies: the Pantherinae and Felinae, with the Acinonychinae subsumed into the latter. Pantherinae includes five Panthera and two Neofelis species, while Felinae includes the other 34 species in 12 genera. [11]
The first cats emerged during the Oligocene about 25 million years ago, with the appearance of Proailurus and Pseudaelurus . The latter species complex was ancestral to two main lines of felids: the cats in the extant subfamilies and a group of extinct "saber-tooth" felids of the subfamily Machairodontinae, which range from the type genus Machairodus of the late Miocene to Smilodon of the Pleistocene. The "false saber-toothed cats", the Barbourofelidae and Nimravidae, are not true cats but are closely related. Together with the Felidae, Viverridae, hyenas and mongooses, they constitute the Feliformia. [7]
All members of the cat family have the following characteristics in common:
The colour, length and density of their fur are very diverse. Fur colour covers the gamut from white to black, and fur patterns from distinctive small spots, and stripes to small blotches and rosettes. Most cat species are born with spotted fur, except the jaguarundi (Herpailurus yagouaroundi), Asian golden cat (Catopuma temminckii) and caracal (Caracal caracal). The spotted fur of lion (Panthera leo) and cougar (Puma concolor) cubs change to uniform fur during their ontogeny. [8] Those living in cold environments have thick fur with long hair, like the snow leopard (Panthera uncia) and the Pallas's cat (Otocolobus manul). [14] Those living in tropical and hot climate zones have short fur. [9] Several species exhibit melanism with all-black individuals. [25]
In the great majority of cat species, the tail is between a third and a half of the body length, although with some exceptions, like the Lynx species and margay (Leopardus wiedii). [9] Cat species vary greatly in body and skull sizes, and weights:
Most cat species have a haploid number of 18 or 19. Central and South American cats have a haploid number of 18, possibly due to the combination of two smaller chromosomes into a larger one. [30]
Most cat species are also induced ovulators, although the margay appears to be a spontaneous ovulator. [15]
Felidae have type IIx muscle fibers three times more powerful than the muscle fibers of human athletes. [31]
The family Felidae is part of the Feliformia, a suborder that diverged probably about 50.6 to 35 million years ago into several families. [32] The Felidae and the Asiatic linsangs are considered a sister group, which split about 35.2 to 31.9 million years ago. [33]
The earliest cats probably appeared about 35 to 28.5 million years ago. Proailurus is the oldest known cat that occurred after the Eocene–Oligocene extinction event about 33.9 million years ago; fossil remains were excavated in France and Mongolia's Hsanda Gol Formation. [7] Fossil occurrences indicate that the Felidae arrived in North America around 18.5 million years ago. This is about 20 million years later than the Ursidae and the Nimravidae, and about 10 million years later than the Canidae. [34]
In the Early Miocene about 20 to 16.6 million years ago, Pseudaelurus lived in Africa. Its fossil jaws were also excavated in geological formations of Europe's Vallesian, Asia's Middle Miocene and North America's late Hemingfordian to late Barstovian epochs. [35]
In the Early or Middle Miocene, the saber-toothed Machairodontinae evolved in Africa and migrated northwards in the Late Miocene. [36] With their large upper canines, they were adapted to prey on large-bodied megaherbivores. [37] [38] Miomachairodus is the oldest known member of this subfamily. Metailurus lived in Africa and Eurasia about 8 to 6 million years ago. Several Paramachaerodus skeletons were found in Spain. Homotherium appeared in Africa, Eurasia and North America around 3.5 million years ago, and Megantereon about 3 million years ago. Smilodon lived in North and South America from about 2.5 million years ago. This subfamily became extinct in the Late Pleistocene. [36]
Results of mitochondrial analysis indicate that the living Felidae species descended from a common ancestor, which originated in Asia in the Late Miocene epoch. They migrated to Africa, Europe and the Americas in the course of at least 10 migration waves during the past ~11 million years. Low sea levels and interglacial and glacial periods facilitated these migrations. [39] Panthera blytheae is the oldest known pantherine cat dated to the late Messinian to early Zanclean ages about 5.95 to 4.1 million years ago. A fossil skull was excavated in 2010 in Zanda County on the Tibetan Plateau. [40] Panthera palaeosinensis from North China probably dates to the Late Miocene or Early Pliocene. The skull of the holotype is similar to that of a lion or leopard. [41] Panthera zdanskyi dates to the Gelasian about 2.55 to 2.16 million years ago. Several fossil skulls and jawbones were excavated in northwestern China. [42] Panthera gombaszoegensis is the earliest known pantherine cat that lived in Europe about 1.95 to 1.77 million years ago. [43]
Living felids fall into eight evolutionary lineages or species clades. [44] [45] Genotyping of the nuclear DNA of all 41 felid species revealed that hybridization between species occurred in the course of evolution within the majority of the eight lineages. [46]
Modelling of felid coat pattern transformations revealed that nearly all patterns evolved from small spots. [47]
Traditionally, five subfamilies had been distinguished within the Felidae based on phenotypical features: the Pantherinae, the Felinae, the Acinonychinae, [10] and the extinct Machairodontinae and Proailurinae. [48] Acinonychinae used to only contain the genus Acinonyx but this genus is now within the Felinae subfamily. [11]
The following cladogram based on Piras et al. (2013) depicts the phylogeny of basal living and extinct groups. [49]
Felidae |
| |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
The phylogenetic relationships of living felids are shown in the following cladogram: [46]
Felidae |
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Panthera is a genus within the family Felidae, it is one of two extant genera in the subfamily Pantherinae, and contains the largest living members of the cat family. There are 5 living species, the tiger, jaguar, lion, leopard and snow leopard and a number of extinct species.
The term "big cat" is typically used to refer to any of the five living members of the genus Panthera, namely the tiger, lion, jaguar, leopard, and snow leopard, as well as the non-pantherine cheetah and cougar.
Pseudaelurus is a prehistoric cat that lived in Europe, Asia and North America in the Miocene between approximately twenty and eight million years ago. It is considered to be a paraphyletic grade ancestral to living felines and pantherines as well as the extinct machairodonts (saber-tooths), and is a successor to Proailurus. It originated from Eurasia and was the first cat to reach North America, when it entered the continent at about 18.5 Ma ending a 'cat-gap' of 7 million years. The slender proportions of the animal, together with its short, viverrid-like legs, suggest that it may have been an agile climber of trees.
Felis is a genus of small and medium-sized cat species native to most of Africa and south of 60° latitude in Europe and Asia to Indochina. The genus includes the domestic cat. The smallest Felis species is the black-footed cat with a head and body length from 38 to 42 cm. The largest is the jungle cat with a head and body length from 62 to 76 cm.
The African golden cat is a wild cat endemic to the rainforests of West and Central Africa. It is threatened due to deforestation and bushmeat hunting and listed as Vulnerable on the IUCN Red List. It is a close relative of both the caracal and the serval. Previously, it was placed in the genus Profelis. Its body size ranges from 61 to 101 cm with a 16 to 46 cm long tail.
The caracal is a medium-sized wild cat native to Africa, the Middle East, Central Asia, and arid areas of Pakistan and northwestern India. It is characterised by a robust build, long legs, a short face, long tufted ears, relatively short tail, and long canine teeth. Its coat is uniformly reddish tan or sandy, while the ventral parts are lighter with small reddish markings. It reaches 40–50 cm (16–20 in) at the shoulder and weighs 8–19 kg (18–42 lb). It was first scientifically described by German naturalist Johann Christian Daniel von Schreber in 1776. Three subspecies are recognised.
The leopard cat is a small wild cat native to continental South, Southeast, and East Asia. Since 2002 it has been listed as Least Concern on the IUCN Red List as it is widely distributed although threatened by habitat loss and hunting in parts of its range.
Prionailurus is a genus of spotted, small wild cats native to Asia. Forests are their preferred habitat; they feed on small mammals, reptiles and birds, and occasionally aquatic wildlife.
Panthera atrox, better known as the American lion, also called the North American lion, or American cave lion, is an extinct pantherine cat. Panthera atrox lived in North America during the Pleistocene epoch, from around 340,000 to 12,800 years ago. The species was initially described by American paleontologist Joseph Leidy in 1853 based on a fragmentary mandible (jawbone) from Mississippi; the species name ('atrox') means "savage" or "cruel". The status of the species is debated, with some mammalogists and paleontologists considering it a distinct species or a subspecies of Panthera leo, which contains living lions. However, novel genetic evidence has shown that it is instead a distinct species derived from the Eurasian cave or steppe lion, evolving after its geographic isolation in North America. Its fossils have been excavated from Alaska to Mexico. It was about 25% larger than the modern lion, making it one of the largest known felids.
Pardofelis is a genus of the cat family Felidae. This genus is defined as including one species native to Southeast Asia: the marbled cat. Two other species, formerly classified to this genus, now belong to the genus Catopuma.
Felinae is a subfamily of the Felidae and comprises the small cats having a bony hyoid, because of which they are able to purr but not roar. Other authors have proposed an alternative definition for this subfamily, as comprising only the living conical-toothed cat genera with two tribes, the Felini and Pantherini, and excluding the extinct sabre-toothed Machairodontinae.
Leopardus is a genus comprising eight species of small cats native to the Americas. This genus is considered the oldest branch of a genetic lineage of small cats in the Americas whose common ancestor crossed the Bering land bridge from Asia to North America in the late Miocene.
The Pantherinae is a subfamily of the Felidae; it was named and first described by Reginald Innes Pocock in 1917 as only including the Panthera species. The Pantherinae genetically diverged from a common ancestor between 9.32 to 4.47 million years ago and 10.67 to 3.76 million years ago.
The African wildcat is a small wildcat species with sandy grey fur, pale vertical stripes on the sides and around the face. It is native to Africa, West and Central Asia, and is distributed to Rajasthan in India and Xinjiang in China. It inhabits a broad variety of landscapes ranging from deserts to savannas, shrublands and grasslands.
Pristifelis is an extinct genus of feline from the late Miocene. It contains a single species, Pristifelis attica. The first fossil skull of P. attica was excavated near Pikermi in Attica, Greece. Fossils were also excavated near the Moldovan city of Taraclia. It was also discovered in Maragheh, northwestern Iran. P. attica was bigger in body size than a European wildcat but probably smaller than a serval. The species was first described as Felis attica by Johann Andreas Wagner in 1857. Due to size differences, it was proposed as type species for the genus Pristifelis proposed in 2012.
Panthera blytheae is an extinct species of pantherine felid that lived during the late Messinian to early Zanclean ages approximately 5.95–4.1 million years ago.The first fossils were excavated in August 2010 in the Zanda Basin located in the Ngari Prefecture on the Tibetan Plateau; they were described and named in 2014.
Caracal is a genus in the subfamily Felinae of the family Felidae. It was proposed by John Edward Gray in 1843 who described a skin from the Cape of Good Hope in the collection of the Natural History Museum, London. Historically, it was considered to be a monotypic genus, consisting of only the type species: the caracal C. caracal.
Miopanthera is an extinct genus of Pseudaelurus-grade felids.
Pachypanthera is an extinct genus of pantherine felid that was recovered from the Late Miocene-aged Khorat sand pits in northeastern Thailand. It contains a single species, Pachypanthera piriyai, named and described in 2023.