Hypanthium

Last updated
Punica grantum.JPG
Young Punica granatum Tunisia 2011.jpg
In a pomegranate flower, Punica granatum , the petals, except for their fused bases, soon fall. The hypanthium with sepal lobes and stamens still attached develops to form the outer layer of the fruit.

In angiosperms, a hypanthium or floral cup [1] [2] [3] is a structure where basal portions of the calyx, the corolla, and the stamens form a cup-shaped tube. It is sometimes called a floral tube, a term that is also used for corolla tube and calyx tube.[ citation needed ] It often contains the nectaries of the plant. It is present in many plant families, although varies in structural dimensions and appearance. [4] This differentiation between the hypanthium in particular species is useful for identification. Some geometric forms are obconic shapes as in toyon, whereas some are saucer-shaped as in Mitella caulescens .

Contents

Its presence is diagnostic of many families, including the Rosaceae, Grossulariaceae, and Fabaceae. In some cases, it can be so deep, with such a narrow top, that the flower can appear to have an inferior ovary - the ovary is below the other attached floral parts. The hypanthium is known by different common names in differing species. In the eucalypts, it is referred to as the gum nut; in roses it is called the hip.

Variations in plant species

Ovary superior to hypanthium Ovary position superior with hypanthium.png
Ovary superior to hypanthium
In Spiraea, the hypanthium supports a nectar-producing "disk" which is ring-shaped and may have lobes as it does here. The stamens arise between the petals and the disk. 2013-04-29 16-35-13-fleur-58f.jpg
In Spiraea , the hypanthium supports a nectar-producing "disk" which is ring-shaped and may have lobes as it does here. The stamens arise between the petals and the disk.
Hypanthium in Rosa Flower morphology - longitudinal section showing achene formation and hypanthium in Rosa-01.svg
Hypanthium in Rosa
Narcissus pseudonarcissus, showing from the upper bend to the tip of the flower: spathe, ovary, hypanthium, tepals, corona 20140226Narcissus2.jpg
Narcissus pseudonarcissus, showing from the upper bend to the tip of the flower: spathe, ovary, hypanthium, tepals, corona

In myrtles, the hypanthium can either surround the ovary loosely or tightly; in some cases, it can be fused to the walls of the ovary. It can vary in length. The rims around the outside of the hypanthium contain the calyx lobes or free sepals, petals and either the stamen or multiple stamen that are attached at one or two points.

The flowers of the family Rosaceae, or the rose family, always have some type of hypanthium or at least a floral cup from which the sepals, petals and stamens all arise, and which is lined with nectar-producing tissue known as nectaries. The nectar is a sugary substance that attracts birds and bees to the flower, who then take the pollen from the lining of the hypanthium and transfer it to the next flower they visit, usually a neighbouring plant. [5]

The stamens borne on the hypanthium are the pollen-producing reproductive organs of the flower. The hypanthium helps in many ways with the reproduction and cross pollination pathways of most plants. It provides weather protection and a medium to sustain the lost pollen, increasing the probability of fertility and cross-pollination. [6] The retained pollen can then attach to pollinators such as birds, bees, moths, beetles, bats, butterflies and other animals. Wind can act as an instigator for fertilisation. The hypanthium is also an adaptive feature for structural support. It helps the stem fuse with the flower, in turn strengthening the bond and overall stability and integrity. [7]

Related Research Articles

<span class="mw-page-title-main">Asteraceae</span> Large family of flowering plants

The family Asteraceae, with the original name Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae were first described in the year 1740. The number of species in Asteraceae is rivaled only by the Orchidaceae, and which is the larger family is unclear as the quantity of extant species in each family is unknown.

<span class="mw-page-title-main">Petal</span> Part of most types of flower

Petals are modified leaves that surround the reproductive parts of flowers. They are often brightly colored or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the corolla. Petals are usually accompanied by another set of modified leaves called sepals, that collectively form the calyx and lie just beneath the corolla. The calyx and the corolla together make up the perianth, the non-reproductive portion of a flower. When the petals and sepals of a flower are difficult to distinguish, they are collectively called tepals. Examples of plants in which the term tepal is appropriate include genera such as Aloe and Tulipa. Conversely, genera such as Rosa and Phaseolus have well-distinguished sepals and petals. When the undifferentiated tepals resemble petals, they are referred to as "petaloid", as in petaloid monocots, orders of monocots with brightly colored tepals. Since they include Liliales, an alternative name is lilioid monocots.

<span class="mw-page-title-main">Stamen</span> Male organ of a flower

The stamen is the pollen-producing reproductive organ of a flower. Collectively, the stamens form the androecium.

<span class="mw-page-title-main">Sepal</span> Any of the separate parts of the calyx of a flower (excluding the bracts), usually green

A sepal is a part of the flower of angiosperms. Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom.

<span class="mw-page-title-main">Geraniaceae</span> Family of plants

Geraniaceae is a family of flowering plants placed in the order Geraniales. The family name is derived from the genus Geranium. The family includes both the genus Geranium and the garden plants called geraniums, which modern botany classifies as genus Pelargonium, along with other related genera.

<span class="mw-page-title-main">Achene</span> Class of simple non-opening dry fruits

An achene, also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate and indehiscent. Achenes contain a single seed that nearly fills the pericarp, but does not adhere to it. In many species, what is called the "seed" is an achene, a fruit containing the seed. The seed-like appearance is owed to the hardening of the fruit wall (pericarp), which encloses the solitary seed so closely as to seem like a seed coat.

<span class="mw-page-title-main">Plant reproductive morphology</span> Parts of plant enabling sexual reproduction

Plant reproductive morphology is the study of the physical form and structure of those parts of plants directly or indirectly concerned with sexual reproduction.

<span class="mw-page-title-main">Commelinaceae</span> Family of flowering plants

Commelinaceae is a family of flowering plants. In less formal contexts, the group is referred to as the dayflower family or spiderwort family. It is one of five families in the order Commelinales and by far the largest of these with about 731 known species in 41 genera. Well known genera include Commelina (dayflowers) and Tradescantia (spiderworts). The family is diverse in both the Old World tropics and the New World tropics, with some genera present in both. The variation in morphology, especially that of the flower and inflorescence, is considered to be exceptionally high amongst the angiosperms.

<span class="mw-page-title-main">Perianth</span> Collective term for the sepals and petals, or either of them if one is absent

The perianth is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when called a perigone. The term perianth is derived from Greek περί and άνθος, while perigonium is derived from περί and γόνος . In the mosses and liverworts (Marchantiophyta), the perianth is the sterile tubelike tissue that surrounds the female reproductive structure.

<span class="mw-page-title-main">Gynoecium</span> Female organs of a flower

Gynoecium is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of pistils and is typically surrounded by the pollen-producing reproductive organs, the stamens, collectively called the androecium. The gynoecium is often referred to as the "female" portion of the flower, although rather than directly producing female gametes, the gynoecium produces megaspores, each of which develops into a female gametophyte which then produces egg cells.

<span class="mw-page-title-main">Nectar</span> Sugar-rich liquid produced by many flowering plants, that attracts pollinators and insects

Nectar is a sugar-rich liquid produced by plants in glands called nectaries or nectarines, either within the flowers with which it attracts pollinating animals, or by extrafloral nectaries, which provide a nutrient source to animal mutualists, which in turn provide herbivore protection. Common nectar-consuming pollinators include mosquitoes, hoverflies, wasps, bees, butterflies and moths, hummingbirds, honeyeaters and bats. Nectar plays a crucial role in the foraging economics and evolution of nectar-eating species; for example, nectar foraging behavior is largely responsible for the divergent evolution of the African honey bee, A. m. scutellata and the western honey bee.

<span class="mw-page-title-main">Calyceraceae</span> Family of flowering plants

Calyceraceae is a plant family in the order Asterales. The natural distribution of the about sixty species belonging to this family is restricted to the southern half of South America. The species of the family resemble both the family Asteraceae and the Dipsacaceae.

<span class="mw-page-title-main">Flower</span> Reproductive structure in flowering plants

A flower, also known as a bloom or blossom, is the reproductive structure found in flowering plants. Flowers consist of a combination of vegetative organs – sepals that enclose and protect the developing flower, petals that attract pollinators, and reproductive organs that produce gametophytes, which in flowering plants produce gametes. The male gametophytes, which produce sperm, are enclosed within pollen grains produced in the anthers. The female gametophytes are contained within the ovules produced in the carpels.

This page provides a glossary of plant morphology. Botanists and other biologists who study plant morphology use a number of different terms to classify and identify plant organs and parts that can be observed using no more than a handheld magnifying lens. This page provides help in understanding the numerous other pages describing plants by their various taxa. The accompanying page—Plant morphology—provides an overview of the science of the external form of plants. There is also an alphabetical list: Glossary of botanical terms. In contrast, this page deals with botanical terms in a systematic manner, with some illustrations, and organized by plant anatomy and function in plant physiology.

Tetrachondra is a plant genus and a member of the family Tetrachondraceae. It comprises two species of creeping succulent, perennial, aquatic or semi-aquatic herbaceous plants. Its distribution range is disjunct: one species is endemic to New Zealand while the other one is endemic to southern Patagonia and Tierra del Fuego. These plants bear essential oils.

This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary of leaf morphology. For other related terms, see Glossary of phytopathology, Glossary of lichen terms, and List of Latin and Greek words commonly used in systematic names.

<span class="mw-page-title-main">Whorl (botany)</span> Arrangement of plant parts around the stem

In botany, a whorl or verticil is a whorled arrangement of leaves, sepals, petals, stamens, or carpels that radiate from a single point and surround or wrap around the stem or stalk. A leaf whorl consists of at least three elements; a pair of opposite leaves is not called a whorl.

<i>Passiflora mixta</i> Species of vine

Passiflora mixta, from the family Passifloraceae is also known as curuba, curuba de indio, curuba de monte, curubita, curuba (Colombia), parcha (Venezuela), and taxo (Ecuador). Originally, it derived from the monophyletic Passiflora subgenus Tacsonia. Passiflora mixta is endemic to the Americas. A perennial vine, the Passiflora mixta is pink to orange-red in color.

<span class="mw-page-title-main">Nectar spur</span>

A nectar spur is a hollow extension of a part of a flower. The spur may arise from various parts of the flower: the sepals, petals, or hypanthium, and often contain tissues that secrete nectar (nectaries). Nectar spurs are present in many clades across the angiosperms, and are often cited as an example of convergent evolution.

<span class="mw-page-title-main">Floral morphology</span> Study of flower structures

In botany, floral morphology is the study of the diversity of forms and structures presented by the flower, which, by definition, is a branch of limited growth that bears the modified leaves responsible for reproduction and protection of the gametes, called floral pieces.

References

  1. Foster 2014, Hypanthium.
  2. Beentje, H.; Williamson, J. (2010). The Kew Plant Glossary: an Illustrated Dictionary of Plant Terms. Royal Botanic Gardens, Kew: Kew Publishing.
  3. Hickey, M.; King, C. (2001). The Cambridge Illustrated Glossary of Botanical Terms. Cambridge University Press.
  4. Cronquist 1981.
  5. Givnish 1997.
  6. Clarke 2004.
  7. Snow 2003.

Bibliography

Books

Websites

Foster, Tony. "Botany Word of the Day". Phytography. Retrieved 27 November 2014.