Aethocrinus Temporal range: | |
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Scientific classification | |
Domain: | Eukaryota |
Kingdom: | Animalia |
Phylum: | Echinodermata |
Class: | Crinoidea |
Family: | † Aethocrinidae |
Genus: | † Aethocrinus |
Aethocrinus is an early-diverging crinoid reported from the Early Ordovician. [1] Its five arms bifurcate.
The contested Echmatocrinus notwithstanding, it is a candidate for the earliest true member of the crinoid class – though not being a member of the crown group, quite where within the stem group to draw the distinction between eocrinoids and crinoids is a somewhat arbitrary decision.
An echinoderm is any member of the phylum Echinodermata. The adults are recognisable by their radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the sea lilies or "stone lilies". Adult echinoderms are found on the sea bed at every ocean depth, from the intertidal zone to the abyssal zone. The phylum contains about 7,000 living species, making it the second-largest grouping of deuterostomes, after the chordates. Echinoderms are the largest entirely marine phylum. The first definitive echinoderms appeared near the start of the Cambrian.
Crinoids are marine invertebrates that make up the class Crinoidea. Crinoids that are attached to the sea bottom by a stalk in their juvenile form are commonly called sea lilies, while the unstalked forms, called feather stars or comatulids, are members of the largest crinoid order, Comatulida. Crinoids are echinoderms in the phylum Echinodermata, which also includes the starfish, brittle stars, sea urchins and sea cucumbers. They live in both shallow water and in depths as great as 9,000 meters (30,000 ft).
A reef is a ridge or shoal of rock, coral or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic (non-living) processes such as deposition of sand or wave erosion planing down rock outcrops. However, reefs such as the coral reefs of tropical waters are formed by biotic (living) processes, dominated by corals and coralline algae. Artificial reefs such as shipwrecks and other man-made underwater structures may occur intentionally or as the result of an accident, and are sometimes designed to increase the physical complexity of featureless sand bottoms to attract a more diverse range of organisms. Reefs are often quite near to the surface, but not all definitions require this.
Crinozoa is a subphylum of mostly sessile echinoderms, of which the crinoids, or sea lilies and feather stars, are the only extant members. Crinozoans have an extremely extensive fossil history, which may or may not extend into the Precambrian.
Pentacrinites is an extinct genus of crinoids that lived from the Hettangian to the Bathonian of Asia, Europe, North America, and New Zealand. Their stems are pentagonal to star-shaped in cross-section and are the most commonly preserved parts. Pentacrinites are commonly found in the Pentacrinites Bed of the Early Jurassic of Lyme Regis, Dorset, England. Pentacrinites can be recognized by the extensions all around the stem, which are long, unbranching, and of increasing length further down, the very small cup and 5 long freely branching arms.
The Decorah Shale is a fossiliferous shale that makes up the lowermost formation in the Galena Group. The Decorah lies above the Platteville Limestone and below the Cummingsville Formation in the sedimentary sequence that formed from the shallow sea that covered central North America during Ordovician Time. The Decorah consists of three members : Spechts Ferry, Guttenberg, and Ion. The Spechts Ferry member is organic-rich and suggests a large influx of terrigenous sediment during deposition. The Guttenberg is characterized by nodular calcareous beds and contains several K-bentonite deposits. The Ion Member, present in the southern Decorah in Iowa, is characterized by alternating beds of shale and limestone.
Pilumnoidea is a superfamily of crabs, whose members were previously included in the Xanthoidea. The three families are unified by the free articulation of all the segments of the male crab's abdomen and by the form of the gonopods. The earliest fossils assigned to this group are of Eocene age.
The Late Silurian to Early Devonian Keyser Formation is a mapped limestone bedrock unit in Pennsylvania, Maryland, Virginia, and West Virginia.
The Mississippian Borden Formation is a mapped bedrock unit in Kentucky, Indiana, Illinois, Ohio, West Virginia, and Tennessee. It has many members, which has led some geologists to consider it a group rather than a formation.
The Brassfield Formation, named by A.F. Foerste in 1906, is a limestone and dolomite formation exposed in Arkansas, Ohio, Kentucky, Indiana, Tennessee and West Virginia in the United States. It is Early Silurian in age and well known for its abundant echinoderms, corals and stromatoporoids. In Ohio, where the unit has escaped dolomitization, the Brassfield is an encrinite biosparite with numerous crinoid species.
Comatulida is an order of crinoids. Members of this order are known as feather stars and mostly do not have a stalk as adults. The oral surface with the mouth is facing upwards and is surrounded by five, often divided rays with feathery pinnules. Comatulids live on the seabed and on reefs in tropical and temperate waters.
Paleontology in New Hampshire refers to paleontological research occurring within or conducted by people from the U.S. state of New Hampshire. Fossils are very rare in New Hampshire because so much of the state's geology is highly metamorphic. The state's complicated geologic history has made dating its rocks and the fossils they contain "a difficult task." The state's Devonian rocks are especially metamorphosed, yet its Mississippian rocks formed too recently to have been subject to the same metamorphism. Nevertheless, despite the geologic complications some fossils have been discovered in the state.
Paleontology in Missouri refers to paleontological research occurring within or conducted by people from the U.S. state of Missouri. The geologic column of Missouri spans all of geologic history from the Precambrian to present with the exception of the Permian, Triassic, and Jurassic. Brachiopods are probably the most common fossils in Missouri.
Bourgueticrinida is an order of crinoids that typically live deep in the ocean. Members of this order are attached to the seabed by a slender stalk and are known as sea lilies. While other groups of crinoids flourished during the Permian, bourgueticrinids along with other extant orders did not appear until the Triassic, following a mass extinction event in which nearly all crinoids died out.
The Camerata or camerate crinoids are an extinct subclass of Paleozoic stalked crinoids. They were some of the earliest crinoids to originate during the Early Ordovician, reached their maximum diversity during the Mississippian, and became extinct during the Permian–Triassic extinction event. Camerates are the sister group of Pentacrinoidea, which contains all other crinoids. The two largest camerate subgroups are the orders Diplobathrida and Monobathrida.
The Ganigobis Formation is a Late Carboniferous (Gzhelian) to Early Permian (Artinskian) geologic formation of the Dwyka Group in the ǁKaras Region of southeastern Namibia and the Northern Cape of South Africa. The widespread formation was deposited in the Aranos and Karoo Basins of southern Africa.
Disparida is an parvclass of extinct marine animals in the class Crinoidea. Disparids are a speciose and morphologically diverse group of crinoids distinguished by their monocyclic calyx and slender arms without pinnules. They range from the Early Ordovician (Tremadocian) to Middle Permian, reaching their highest diversity during the Late Ordovician.
Pentacrinoidea is a subclass of crinoids containing all members of Crinoidea except for the exclusively Paleozoic camerates. It was originally named in 1918 by Otto Jaekel, who hypothesized a fundamental split between camerate and non-camerate crinoids. Later workers doubted this interpretation, and Pentacrinoidea was rarely used during the rest of the 20th century. Recent phylogenetic work has provided strong support for Jaekel's hypothesis, and Pentacrinoidea was reinstated in a 2017 revision of crinoid systematics.
Flexibilia is a superorder of specialized Paleozoic crinoids. They exhibited a conserved body plan and consistent suite of characteristics throughout their long history. Previously considered a subclass with unclear affinities, later investigation determined that flexibles are well-nested within Cladida, a broad group ancestral to living articulate crinoids. The Ordovician cladid Cupulocrinus acts as an intermediate form linking the generalized anatomy of other early cladids with the distinctive anatomy of flexibles, and several studies have considered it to be ancestral to the rest of the group. Although flexibles never reached the same abundance or diversity as many other crinoid groups, they remained a reliable component of crinoid faunas, particularly from the Silurian onwards. Flexible fossils are very rare in the Ordovician, but the Late Ordovician appears to have been an interval of rapid diversification for the group.