Agmata Temporal range: | |
---|---|
Volborthella | |
Scientific classification | |
Kingdom: | Animalia |
Phylum: | † Agmata Yochelson, 1977 [1] |
Taxa | |
Volborthellida |
Agmata is a proposed extinct phylum of small animals with a calcareous conical shell. They were originally thought to be cephalopods or annelid worms. The living animals filled up to five-sixths of their shell with laminae, angled layers composed of grains of quartz or calcium carbonate detritus from the environment cemented together, with larger grains near the shell wall and smaller grains near the center. A very fine tube ran through the center of the shell. The grains may be of quartz or calcium carbonate, but are of specific shapes and materials that are rare in the surrounding rock. Though the body of the living animal is not preserved, it had to be able to find, choose, and retrieve rare grains from its environment to build the laminae.
The phylum's name comes from the Greek word for "fragments", referring to these fine fragments and grains of detritus. [1] It was proposed by the paleontologist and geologist Ellis L. Yochelson (1928–2006) in 1977 to house the agglutinating Early Cambrian fossils Salterella and Volborthella , with the Middle Cambrian Ellisell yochelsoni later included. [2] The poorly known Middle Cambrian fossil Vologdinella was also considered for inclusion, as it has superficial resemblance to the Agmata, but was later excluded from the group. [3]
Currently, the phylum contains only one family, Salterellidae; a second family, Volborthellidae, was originally included but later became a synonym of the former. No orders, classes or superfamilies are used within the phylum, despite the order "Volborthellida" being previously proposed for Volborthella before the phylum's own proposal. The reasoning for this was that taxa of these ranks were not seen as necessary in a phylum with very few genera. [1] [3]
The genera within the group are clearly different: Salterella had a pointed shell made of calcium carbonate with a thin outer layer and a thick inner layer, and secreted a calcium carbonate cement to hold its grains in place. Volborthella was the older genus, had a blunter shell with a shallower opening, and cemented grains in place with organic material that also may have formed the outside surface of the shell. [4]
Fossils are found in large numbers in some areas. Paleontologists have offered several different ideas of how these animals lived: filter-feeding with the points of the shells embedded in the substrate, grazing actively like snails, or lying on their sides on the substrate. Volborthella is found in silt and clay deposits, and apparently lived on tidal mudflats. Attempts to reconcile these genera as members of any other group have been rejected due to basic differences in structure, but not all paleontologists accept them as a phylum; Jones (2007) considers the shell an agglutinating test parallel to that of foraminifers. Yochelson considered Agmata to be complex multicellular animals.
Limestone is a type of carbonate sedimentary rock which is the main source of the material lime. It is composed mostly of the minerals calcite and aragonite, which are different crystal forms of CaCO3. Limestone forms when these minerals precipitate out of water containing dissolved calcium. This can take place through both biological and nonbiological processes, though biological processes, such as the accumulation of corals and shells in the sea, have likely been more important for the last 540 million years. Limestone often contains fossils which provide scientists with information on ancient environments and on the evolution of life.
The cloudinids, an early metazoan family containing the genera Acuticocloudina, Cloudina and Conotubus, lived in the late Ediacaran period about 550 million years ago. and became extinct at the base of the Cambrian. They formed millimetre-scale conical fossils consisting of calcareous cones nested within one another; the appearance of the organism itself remains unknown. The name Cloudina honors the 20th-century geologist and paleontologist Preston Cloud.
Archaeocyatha is a taxon of extinct, sessile, reef-building marine sponges that lived in warm tropical and subtropical waters during the Cambrian Period. It is believed that the centre of the Archaeocyatha origin is now located in East Siberia, where they are first known from the beginning of the Tommotian Age of the Cambrian, 525 million years ago (mya). In other regions of the world, they appeared much later, during the Atdabanian, and quickly diversified into over a hundred families. They became the planet's very first reef-building animals and are an index fossil for the Lower Cambrian worldwide.
Sedimentary rocks are types of rock that are formed by the accumulation or deposition of mineral or organic particles at Earth's surface, followed by cementation. Sedimentation is the collective name for processes that cause these particles to settle in place. The particles that form a sedimentary rock are called sediment, and may be composed of geological detritus (minerals) or biological detritus. The geological detritus originated from weathering and erosion of existing rocks, or from the solidification of molten lava blobs erupted by volcanoes. The geological detritus is transported to the place of deposition by water, wind, ice or mass movement, which are called agents of denudation. Biological detritus was formed by bodies and parts of dead aquatic organisms, as well as their fecal mass, suspended in water and slowly piling up on the floor of water bodies. Sedimentation may also occur as dissolved minerals precipitate from water solution.
An exoskeleton is the external skeleton that supports and protects an animal's body, in contrast to the internal skeleton (endoskeleton) of, for example, a human. In usage, some of the larger kinds of exoskeletons are known as "shells". Examples of exoskeletons within animals include the arthropod exoskeleton shared by chelicerates, myriapods, crustaceans, and insects, as well as the shell of certain sponges and the mollusc shell shared by snails, clams, tusk shells, chitons and nautilus. Some animals, such as the turtle, have both an endoskeleton and an exoskeleton.
Ooids are small, spheroidal, "coated" (layered) sedimentary grains, usually composed of calcium carbonate, but sometimes made up of iron- or phosphate-based minerals. Ooids usually form on the sea floor, most commonly in shallow tropical seas. After being buried under additional sediment, these ooid grains can be cemented together to form a sedimentary rock called an oolite. Oolites usually consist of calcium carbonate; these belong to the limestone rock family. Pisoids are similar to ooids, but are larger than 2 mm in diameter, often considerably larger, as with the pisoids in the hot springs at Carlsbad in the Czech Republic.
Shelly limestone is a highly fossiliferous limestone, composed of a number of fossilized organisms such as brachiopods, bryozoans, crinoids, sponges, corals and mollusks. It varies in color, texture and hardness. Coquina is a poorly indurated form of shelly limestone.
The molluscshell is typically a calcareous exoskeleton which encloses, supports and protects the soft parts of an animal in the phylum Mollusca, which includes snails, clams, tusk shells, and several other classes. Not all shelled molluscs live in the sea; many live on the land and in freshwater.
The small shelly fauna, small shelly fossils (SSF), or early skeletal fossils (ESF) are mineralized fossils, many only a few millimetres long, with a nearly continuous record from the latest stages of the Ediacaran to the end of the Early Cambrian Period. They are very diverse, and there is no formal definition of "small shelly fauna" or "small shelly fossils". Almost all are from earlier rocks than more familiar fossils such as trilobites. Since most SSFs were preserved by being covered quickly with phosphate and this method of preservation is mainly limited to the late Ediacaran and early Cambrian periods, the animals that made them may actually have arisen earlier and persisted after this time span.
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks. Around 85,000 extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied.
Brachiopods, phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a stalk-like pedicle projects from an opening near the hinge of one of the valves, known as the pedicle or ventral valve. The pedicle, when present, keeps the animal anchored to the seabed but clear of sediment which would obstruct the opening.
Volborthella is an animal of uncertain classification, whose fossils pre-date 530 million years ago. It has been considered for a period a cephalopod. However discoveries of more detailed fossils showed that Volborthella’s small, conical shell was not secreted but built from grains of the mineral silicon dioxide (silica), and that it was not divided into a series of compartments by septa as those of fossil shelled cephalopods and the living Nautilus are. This illusion was a result of the laminated texture of the organisms' tests. Therefore, Volborthella’s classification is now uncertain. It has been speculated that it may in fact represent a sclerite of a larger organism, on the basis of one specimen; however, it may be premature to accept this hypothesis, as the arrangement of sclerites producing this impression may have occurred by chance. The Ordovician scleritome-bearing Curviconophorus, as well as the Halwaxiids, lobopods and echinoderms, demonstrate the diversity of organisms which may produce a scleritome of this nature. The related Campitius was originally suggested to be part of a radula rather than a scleritome, but is now considered a synonym of Volborthella.
Fordilla is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early Cambrian fossils found in North America, Greenland, Europe, the Middle East, and Asia. The genus currently contains three described species, Fordilla germanica, Fordilla sibirica, and the type species Fordilla troyensis.
Salterella is an enigmatic Cambrian genus with a small, conical, calcareous shell that appears to be septate, but is rather filled with stratified laminar deposits. The shell contains grains of sediment, which are obtained selectively by a manner also observed in foramanifera. The genus was established by Elkanah Billings in 1861, and was named after the English palaeontologist John William Salter.
Vologdinella is a poorly known genus of extinct animals of uncertain classification with small cylindrical shells. The animals are known from Middle Cambrian fossils from a Paleozoic limestone in the Chingiz Mountains of Kazakhstan. The genus was established by Russian paleontologist Zakhar Grigoryevich Balashov in 1962 for a single species, Vologdinella antiqua, which was originally described and illustrated as Orthoceras? antiquus by Aleksandr Grigoryevich Vologdin in 1931.
Pojetaia is an extinct genus of early bivalves, one of two genera in the extinct family Fordillidae. The genus is known solely from Early to Middle Cambrian fossils found in North America, Greenland, Europe, North Africa, Asia, and Australia. The genus currently contains two accepted species, Pojetaia runnegari, the type species, and Pojetaia sarhroensis, though up to seven species have been proposed. The genera Buluniella, Jellia, and Oryzoconcha are all considered synonyms of Pojetaia.
Fordillidae is an extinct family of early bivalves and one of two families in the extinct superfamily Fordilloidea. The family is known from fossils of early to middle Cambrian age found in North America, Greenland, Europe, the Middle East, Asia, and Australia. The family currently contains two genera, Fordilla and Pojetaia, each with up to three described species. Due to the size and age of the fossil specimens, Fordillidae species are included as part of the Turkish Small shelly fauna.
Salterellidae is a family of enigmatic fossil genera from the Early to Middle Cambrian. It was originally created for the genus Salterella by Charles Doolittle Walcott, who placed it in the group Pteropoda. It was later placed in Agmata, a proposed extinct phylum by Ellis L. Yochelson which is accepted by some other authors.
Ellisell is a Middle Cambrian genus of fossils from Denmark. It contains only one species, Ellisell yochelsoni. Both the genus and species are named after the paleontologist and geologist Ellis L. Yochelson (1928–2006), who had turned 60 at the time the fossils were first described. The genus was originally placed in the family Salterellidae of the phylum Agmata; this placement was rejected by Yochelson & Kisselev (2003), but was restored by Peel (2016). Ellisell is distinguished from Salterella by its slowly expanding conch and the resulting cylindrical apertural cavity, compared to the latter's more rapidly expanding conch and cone-shaped apertural cavity.
Foraminiferal tests are the tests of Foraminifera.