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The subphylum Chelicerata constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids, as well as a number of extinct lineages, such as the eurypterids and chasmataspidids.
Remipedia is a class of blind crustaceans, closely related to hexapods, found in coastal aquifers which contain saline groundwater, with populations identified in almost every ocean basin so far explored, including in Australia, the Caribbean Sea, and the Atlantic Ocean. The first described remipede was the fossil Tesnusocaris goldichi. Since 1979, at least seventeen living species have been identified in subtropical regions around the world.
Myriapods are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial.
Anomura is a group of decapod crustaceans, including hermit crabs and others. Although the names of many anomurans include the word crab, all true crabs are in the sister group to the Anomura, the Brachyura.
Eucarida is a superorder of the Malacostraca, a class of the crustacean subphylum, comprising the decapods, krill, and Angustidontida. They are characterised by having the carapace fused to all thoracic segments, and by the possession of stalked eyes.
Arachnomorpha is a proposed subdivision or clade of Arthropoda, comprising the group formed by the trilobites and their close relatives (Artiopoda), Megacheira and chelicerates. Under this proposed classification scheme, Arachnomorpha is considered the sister group to Mandibulata.
Pancrustacea is the clade that comprises all crustaceans, including hexapods. This grouping is contrary to the Atelocerata hypothesis, in which Hexapoda and Myriapoda are sister taxa, and Crustacea are only more distantly related. As of 2010, the Pancrustacea taxon was considered well accepted, with most studies recovering Hexapoda within Crustacea. The clade has also been called Tetraconata, referring to having four cone cells in the ommatidia. This name is preferred by some scientists as a means of avoiding confusion with the use of "pan-" to indicate a clade that includes a crown group and all of its stem group representatives.
Terrestrial animals are animals that live predominantly or entirely on land, as compared with aquatic animals, which live predominantly or entirely in the water, and amphibians, which rely on aquatic and terrestrial habitats. Some groups of insects are terrestrial, such as ants, butterflies, earwigs, cockroaches, grasshoppers and many others, while other groups are partially aquatic, such as mosquitoes and dragonflies, which pass their larval stages in water.
Mandibulata, is one of two major clades of living arthropods alongside Chelicerata. It comprises the extant groups Myriapoda and Pancrustacea. The name "Mandibulata" refers to the mandibles, a modified pair of limbs used in food processing, the presence of which are characteristic of most members of the group. Members of the group are referred to as mandibulates.
The (pan)arthropod head problem is a long-standing zoological dispute concerning the segmental composition of the heads of the various arthropod groups, and how they are evolutionarily related to each other. While the dispute has historically centered on the exact make-up of the insect head, it has been widened to include other living arthropods, such as chelicerates, myriapods, and crustaceans, as well as fossil forms, such as the many arthropods known from exceptionally preserved Cambrian faunas. While the topic has classically been based on insect embryology, in recent years a great deal of developmental molecular data has become available. Dozens of more or less distinct solutions to the problem, dating back to at least 1897, have been published, including several in the 2000s.
The Myriochelata or Paradoxopoda, is a proposed grouping of arthropods comprising the Myriapoda and Chelicerata. If this proposition holds true, the Myriochelata are the sister clade to the Pancrustacea, comprising classic crustaceans and hexapods.
Apposition eyes are the most common form of eye, and are presumably the ancestral form of compound eye. They are found in all arthropod groups, although they may have evolved more than once within this phylum. Some annelids and bivalves also have apposition eyes. They are also possessed by Limulus, the horseshoe crab, and there are suggestions that other chelicerates developed their simple eyes by reduction from a compound starting point. Some caterpillars appear to have evolved compound eyes from simple eyes in the opposite fashion.
Arthropods are invertebrate animals in the phylum Arthropoda. They possess an exoskeleton with a cuticle made of chitin, often mineralised with calcium carbonate, a metamerically segmented body, and paired jointed appendages. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. They are an extremely diverse group, with up to 10 million species.
Xenocarida is a proposed clade inside the subphylum Crustacea that comprises two classes that were discovered in the 20th century: Remipedia and Cephalocarida. Both groups are marine hermaphrodites. The clade was recovered as the sister groups to Hexapoda.
The Articulata hypothesis is the grouping in a higher taxon of animals with segmented bodies, consisting of Annelida and Panarthropoda. This theory states that these groups are descended from a common segmented ancestor. The Articulata hypothesis is an alternative to the hypothesis that ecdysis is a primitive characteristic – this would place Panarthropoda in the group Ecdysozoa.
Crustaceomorpha is a proposed clade of arthropods that includes crustaceans and numerous extinct groups. Synapomorphies for the clade are that the larval antenna is a feeding or locomotory organ, and there are six endopodal podomeres in post-antennal limbs.
Wingertshellicus is an extinct genus of arthropod that has been found in Hunsrück Slate, that is located in the Rhenish Massif in Germany, and lived about 405 million years ago, during the Lower Emsian.
The subphylum Hexapoda or hexapods comprises the largest clade of arthropods and includes most of the extant arthropod species. It includes the crown group Insecta, as well as three much smaller groups of wingless arthropods that were once considered insects: Collembola (springtails), Protura (coneheads) and Diplura. The insects and springtails are very abundant and are some of the most important pollinators, basal consumers, scavengers/detritivores and micropredators in terrestrial environments.
Crustaceans belong to the subphylum Crustacea, and form a large, diverse group of arthropods including decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, barnacles, copepods, opossum shrimps, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. The three classes Cephalocarida, Branchiopoda and Remipedia are more closely related to the hexapods than they are to any of the other crustaceans.
Antennulata is a proposed clade of arthropods comprising Artiopoda, Marrellomorpha and Mandibulata. As its name suggests, a proposed unifying feature of the clade is the presence of antennulae. It contrasts with the opposing Arachnomorpha hypothesis, which instead posits that artiopods and marrellomorphs are more closely related to Chelicerates.
References
- 1 2 3 4 Koenemann, Stefan; Jenner, Ronald A.; Hoenemann, Mario; Stemme, Torben; von Reumont, Björn M. (2010). "Arthropod phylogeny revisited, with a focus on crustacean relationships" (PDF). Arthropod Structure & Development. 39 (2–3): 88–110. doi:10.1016/j.asd.2009.10.003. PMID 19854296.
- ↑ Richard Heymons (1901). "Die Entwicklungsgeschichte der Scolopender". Zoologica . Stuttgart: Erwin Nägele. 33: 1–244, tables i–viii. doi: 10.5962/bhl.title.1587 .
- ↑ Cook, C. E; Yue, Q.; Akam, M. (2005). "Mitochondrial genomes suggest that hexapods and crustaceans are mutually paraphyletic". Proceedings of the Royal Society B: Biological Sciences. 272 (1569): 1295–1304. doi:10.1098/rspb.2004.3042. PMC 1564108 . PMID 16024395.
- ↑ Guillaume Lecointre & Hervé Le Guyader (2006). The Tree of Life: A Phylogenetic Classification . Harvard University Press. ISBN 978-0-674-02183-9.
- ↑ Maximilian J. Telford & Richard H. Thomas (1995). "Demise of the Atelocerata?". Nature . 376 (6536): 123–124. Bibcode:1995Natur.376..123T. doi: 10.1038/376123a0 .
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