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The Mississippian is a subperiod in the geologic timescale or a subsystem of the geologic record. It is the earlier of two subperiods of the Carboniferous period lasting from roughly 358.9 to 323.2 million years ago. As with most other geochronologic units, the rock beds that define the Mississippian are well identified, but the exact start and end dates are uncertain by a few million years. The Mississippian is so named because rocks with this age are exposed in the Mississippi Valley.
Xenophyophorea is a clade of foraminiferans. Xenophyophores are multinucleate unicellular organisms found on the ocean floor throughout the world's oceans, at depths of 500 to 10,600 metres. They are a kind of foraminiferan that extract minerals from their surroundings and use them to form an exoskeleton known as a test.
Foraminifera are single-celled organisms, members of a phylum or class of Rhizarian protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an external shell of diverse forms and materials. Tests of chitin are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within the seafloor sediment, while a smaller number float in the water column at various depths, which belong to the suborder Globigerinina. Fewer are known from freshwater or brackish conditions, and some very few (nonaquatic) soil species have been identified through molecular analysis of small subunit ribosomal DNA.
The Rhizaria are a diverse and species-rich supergroup of mostly unicellular eukaryotes. Except for the Chlorarachniophytes and three species in the genus Paulinella in the phylum Cercozoa, they are all non-photosynthethic, but many foraminifera and radiolaria have a symbiotic relationship with unicellular algae. A multicellular form, Guttulinopsis vulgaris, a cellular slime mold, has been described. This group was used by Cavalier-Smith in 2002, although the term "Rhizaria" had been long used for clades within the currently recognized taxon. Being described mainly from rDNA sequences, they vary considerably in form, having no clear morphological distinctive characters (synapomorphies), but for the most part they are amoeboids with filose, reticulose, or microtubule-supported pseudopods. In the absence of an apomorphy, the group is ill-defined, and its composition has been very fluid. Some Rhizaria possess mineral exoskeletons, which are in different clades within Rhizaria made out of opal, celestite, or calcite. Certain species can attain sizes of more than a centimeter with some species being able to form cylindrical colonies approximately 1 cm in diameter and greater than 1 m in length. They feed by capturing and engulfing prey with the extensions of their pseudopodia; forms that are symbiotic with unicellular algae contribute significantly to the total primary production of the ocean.
The Fusulinida is an extinct order within the Foraminifera in which the tests are traditionally considered to have been composed of microgranular calcite. Like all forams, they were single-celled organisms. In advanced forms the test wall was differentiated into two or more layers. Loeblich and Tappan, 1988, gives a range from the Lower Silurian to the Upper Permian, with the fusulinid foraminifera going extinct with the Permian–Triassic extinction event. While the latter is true, a more supported projected timespan is from the Mid-Carboniferous period.
The Rotaliida are an order of Foraminifera, characterized by multilocular tests (shells) composed of bilamellar perforate hyaline lamellar calcite that may be optically radial or granular.
The Textulariida are an order of foraminifera that produce agglutinated shells or tests. An agglutinated test is one made of foreign particles glued together with an organic or calcareous cement to form an external shell on the outside of the organism. Commonly, the order had been made up of all species of Foraminifera with these types of shells, but genetic studies indicate these organisms do not form an evolutionary group, and several superfamilies in the order have been moved to the order Allogromiida. The remaining forms are sometimes divided into three orders: the Trochamminida and Lituolida, which have organic cement, and the Textulariida sensu stricto, which use a calcareous cement. All three orders or superfamilies are known as fossils from the Cambrian onwards.
Carterinida is an order of multi-chambered foraminifera within the Globothalamea. Members of this order form hard tests out of thin calcite rods known as spicules, which are held together by a proteinaceous matrix. As of August 2023, the order contains a single family, Carterinidae.
The Spirillinida are an order of foraminifera in which the test, or shell, primitively consists of an enrolled open tube, coming after the proloculus, wound planospirally or conically, commonly composed of an optically single crystal of calcite. The aperture is a simple opening at the end of the tube. Advanced forms with more than one chamber may consist of a few crystals, or rarely, a mosaic of crystals of calcite.
The Robertinida are an order of Middle Triassic to recent, benthic foraminifera in which tests are planispirally to trochospirally coiled, with chambers provided with an internal partition and hyaline perforated walls composed of optically radiated aragonite.
Cibicides is a genus of cosmopolitan benthic foraminifera known from at least as far back as the Paleocene that extends down to the present.
Discorbis is a genus of benthic Foraminifera, that made its first appearance during the Eocene. Its present distribution is cosmopolitan.
Globigerinoides is an extant genus of shallow-water planktonic foraminifera of family Globigerinidae. First appearing in the Oligocene these foraminifera are found in all modern oceans. Species of this genus occupy the euphotic zone, generally at depths between 10-50m, in waters which cover a range of salinities and temperatures. They are a shorter lived species, especially when compared to Globorotalia genus. As a genus Globigerinoides is widely used in various fields of research including biostratigraphy, isotope geochemistry, biogeochemistry, climatology, and oceanography.
Helen Niña Tappan Loeblich was an American micropaleontologist who was a professor of geology at the University of California, Los Angeles, a United States Geological Survey (USGS) biostratigrapher, and a scientific illustrator whose micropaleontology specialty was research on Cretaceous foraminifera.
Alfred R. Loeblich Jr (1914–1994) was an American micropaleontologist. He was married to Helen Niña Tappan Loeblich and the two co-authored a number of important works on the Foraminifera and related organisms.
Tubothalamea is a taxonomic class established for foraminiferans with tubular chambers. Includes the porcelaceous and agglutinated Miliolida and the monocrystalline and agglutinated Spirillinida. It is one of two classes of multichambered foraminifera based on SSU rDNA molecular studies with consideration of major morphological trands, the other being the Globothalamea.
Globothalamea comprises a class of multichambered foraminifera based in part on SSU rDNA evidence; the other is Tubothalamea.
"Monothalamea" is a grouping of foraminiferans, traditionally consisting of all foraminifera with single-chambered tests. Recent work has shown that the grouping is paraphyletic, and as such does not constitute a natural group; nonetheless, the name "monothalamea" continues to be used by foraminifera workers out of convenience.
Bruce William Hayward is a New Zealand geologist, marine ecologist, and author. He is known as a leading expert on living and fossil foraminifera.
Foraminiferal tests are the tests of Foraminifera.
References
- ↑ "Foraminifera - The World Foraminifera Database - Baggina Cushman, 1926". www.marinespecies.org. Retrieved 2021-10-12.
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