Foraminifera are single-celled organisms, members of a phylum or class of Rhizarian protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly an external shell of diverse forms and materials. Tests of chitin are believed to be the most primitive type. Most foraminifera are marine, the majority of which live on or within the seafloor sediment, while a smaller number float in the water column at various depths, which belong to the suborder Globigerinina. Fewer are known from freshwater or brackish conditions, and some very few (nonaquatic) soil species have been identified through molecular analysis of small subunit ribosomal DNA.
The Globigerinina is a suborder of foraminiferans that are found as marine plankton. They produce hyaline calcareous tests, and are known as fossils from the Jurassic period onwards. The group has included more than 100 genera and over 400 species, of which about 30 species are extant. One of the most important genera is Globigerina; vast areas of the ocean floor are covered with Globigerina ooze, dominated by the shells of planktonic forms.
The Allogromiida is an order of single-chambered, mostly organic-walled foraminiferans, including some that produce agglutinated tests (Lagynacea). Genetic studies indicate that some foraminiferans with agglutinated tests, previously included in the Textulariida or as their own order Astrorhizida, may also belong here. Allogromiids produce relatively simple tests, usually with a single chamber, similar to those of other protists such as Gromia. They are found as both marine and freshwater forms, and are the oldest forms known from the fossil record.
The Fusulinida is an extinct order within the Foraminifera in which the tests are traditionally considered to have been composed of microgranular calcite. Like all forams, they were single-celled organisms. In advanced forms the test wall was differentiated into two or more layers. Loeblich and Tappan, 1988, gives a range from the Lower Silurian to the Upper Permian, with the fusulinid foraminifera going extinct with the Permian–Triassic extinction event. While the latter is true, a more supported projected timespan is from the Mid-Carboniferous period.
The Rotaliida are an order of Foraminifera, characterized by multilocular tests (shells) composed of bilamellar perforate hyaline lamellar calcite that may be optically radial or granular.
In biology, a test is the hard shell of some spherical marine animals and protists, notably sea urchins and microorganisms such as testate foraminiferans, radiolarians, and testate amoebae. The term is also applied to the covering of scale insects. The related Latin term testa is used for the hard seed coat of plant seeds.
Carterina is a genus in the family Trochamminidae, composing its own subfamily Carterininae. The genus is described from specimens gathered during the Challenger expedition's circumnavigation of the Earth from 1872-1876.
Carterinida is an order of multi-chambered foraminifera within the Globothalamea. Members of this order form hard tests out of thin calcite rods known as spicules, which are held together by a proteinaceous matrix. As of August 2023, the order contains a single family, Carterinidae.
The Asterigerinacea is a superfamily of Foraminifera included in the order Rotaliida, proposed by Loeblich and Tappan in 1988.
The Spirillinida are an order of foraminifera in which the test, or shell, primitively consists of an enrolled open tube, coming after the proloculus, wound planospirally or conically, commonly composed of an optically single crystal of calcite. The aperture is a simple opening at the end of the tube. Advanced forms with more than one chamber may consist of a few crystals, or rarely, a mosaic of crystals of calcite.
Acervulinacea is a superfamily in the Foraminifera order Rotaliida. The Acervulinacea may be free, or able to move about, or their tests may be attached to some substrate. The early growth stage is spiral, followed by irregular chambers that form an irregular mass, disc, or branching structure. The test wall is of hyaline (glassy) optically radial calcite and is coarsely perforate. Apertures are present only as mural pores.
Lagenida is an order of benthic foraminiferal rhizaria in which the tests (shells) are monolamellar, with walls composed of optically and ultra-structurally radiate calcite, with the crystallographic c-axes perpendicular to the surface. Lagenids first appear in the Upper Silurian and continue to the Recent. They are currently divided into two superfamilies, the older Robuloidacea which range from the Upper Silurian to the Lower Cretaceous (Albian) and the younger Nodosariacea, ranging from the Permian to Recent.
The Robertinida are an order of Middle Triassic to recent, benthic foraminifera in which tests are planispirally to trochospirally coiled, with chambers provided with an internal partition and hyaline perforated walls composed of optically radiated aragonite.
The Cassidulinacea is a superfamily of benthic amoeboid foraminifera in the order Rotaliida that has been extant from the Paleocene down to the present. Tests are composed of secreted, optically radial or granular, perforate calcite with chambers biserially coiled at least in the early part, Apertures are usually an interiomarginal slit, but may become terminal and may have secondary features.
Rotalidia comprises a class of Foraminifera where Foraminifera is regarded as a phylum, that unites Foraminifera that have tests composed of secreted lamellar calcium carbonate, optically radial or granular calcite, or aragonite; separating them from those with porcelaneous, agglutinated, or microgranular, tests, or tests composed of organic compounds. Seven orders are included, the:
Globigerinoides is an extant genus of shallow-water planktonic foraminifera of family Globigerinidae. First appearing in the Oligocene these foraminifera are found in all modern oceans. Species of this genus occupy the euphotic zone, generally at depths between 10-50m, in waters which cover a range of salinities and temperatures. They are a shorter lived species, especially when compared to Globorotalia genus. As a genus Globigerinoides is widely used in various fields of research including biostratigraphy, isotope geochemistry, biogeochemistry, climatology, and oceanography.
Rotaliana is a subclass of benthic Foraminifera with multichambered tests of perforate hyaline calcite. Tests may be planospiral, low or high trochospiral, or serial. Interiors may be complex with secondary chambers and interconnecting canal systems. Rotaliana are separate from the planktonic Globigerinana although both have tests of similar composition. The Textulariana, which contains forms that are rather similar, differs in be agglutinated.
The Spirillinata are a group of Foraminifera established by Maslakova, 1990, for spirally wound forms, where the Foraminifera are regarded as a phylum. Two subclasses are included, the agglutinated Ammodiscana and the calcareous Spirillinana.
Chilostomelloidea is a superfamily of foraminifera in the order Rotaliida. They are found in sediments of Early Cretaceous (Barremian) to the present.
Foraminiferal tests are the tests of Foraminifera.
