| Granofilosea | |
|---|---|
 | |
| Clathrulina elegans | |
Scientific classification  | |
| Domain: | Eukaryota |
| Clade: | Diaphoretickes |
| Clade: | SAR |
| Phylum: | Cercozoa |
| Subphylum: | Reticulofilosa |
| Class: | Granofilosea Cavalier-Smith & Bass 2009 [1] |
| Orders | |
Granofilosea is a class of cercozoan protists in the subphylum Reticulofilosa. [2] Out of the three groups that were traditionally considered heliozoans: the heliomonads, gymnosphaerids and desmothoracids, the latter were recently grouped into this new class.
The following cladogram shows the results of a 2011 phylogenetic analysis: [1]
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Order Desmothoracida, the desmothoracids, are a group of heliozoan protists, usually sessile and found in freshwater environments. The adult is a spherical cell around 10-20 μm in diameter surrounded by a perforated organic lorica, or shell, with many radial pseudopods projecting through the holes to capture food. These are supported by small bundles of microtubules that arise near a point on the nuclear membrane. Unlike other heliozoans, the microtubules are not in any regular geometric array, there does not appear to be a microtubule organizing center, and there is no distinction between the outer and inner cytoplasm.
The centrohelids or centroheliozoa are a large group of heliozoan protists. They include both mobile and sessile forms, found in freshwater and marine environments, especially at some depth.
Cercozoa is a phylum of diverse single-celled eukaryotes. They lack shared morphological characteristics at the microscopic level, and are instead united by molecular phylogenies of rRNA and actin or polyubiquitin. They were the first major eukaryotic group to be recognized mainly through molecular phylogenies. They are the natural predators of many species of bacteria. They are closely related to the phylum Retaria, comprising amoeboids that usually have complex shells, and together form a supergroup called Rhizaria.
Labyrinthulomycetes (ICBN) or Labyrinthulea (ICZN) is a class of protists that produce a network of filaments or tubes, which serve as tracks for the cells to glide along and absorb nutrients for them. The two main groups are the labyrinthulids and thraustochytrids. They are mostly marine, commonly found as parasites on algae and seagrasses or as decomposers on dead plant material. They also include some parasites of marine invertebrates and mixotrophic species that live in a symbiotic relationship with zoochlorella.
Cercomonads are small amoeboflagellates, widespread in aqueous habitats and common in soils.
The Phytomyxea are a class of parasites that are cosmopolitan, obligate biotrophic protist parasites of plants, diatoms, oomycetes and brown algae. They are divided into the orders Plasmodiophorida and Phagomyxida. Plasmodiophorids are best known as pathogens or vectors for viruses of arable crops.
The tectofilosids are a group of filose amoebae with shells. These are composed of organic materials and sometimes collected debris, in contrast to the euglyphids, which produce shells from siliceous scales. The shell usually has a single opening, but in Amphitrema and a few other genera it has two on opposite ends. The cell itself occupies most of the shell. They are most often found on marsh plants such as Sphagnum.
Reticulosida is an order of Endomyxa that was created by Cavalier-Smith in 2003, but subsequently emended in by Bass et al. in 2009 to include only one monotypic family, the Filoretidae.
Discosea is a class of Amoebozoa, consisting of naked amoebae with a flattened, discoid body shape. Members of the group do not produce tubular or subcylindrical pseudopodia, like amoebae of the class Tubulinea. When a discosean is in motion, a transparent layer called hyaloplasm forms at the leading edge of the cell. In some discoseans, short "subpseudopodia" may be extended from this hyaloplasm, but the granular contents of the cell do not flow into these, as in true pseudopodia. Discosean amoebae lack hard shells, but some, like Cochliopodium and Korotnevella secrete intricate organic scales which may cover the upper (dorsal) surface of the cell. No species have flagella or flagellated stages of life.
Monadofilosa is a grouping of Cercozoa. These organisms are single-celled amoeboid protists.
Endomyxa is a group of eukaryotic organisms in the supergroup Rhizaria. They were initially a subphylum of Cercozoa and later a subphylum of Retaria, but several analyses have proven they are a phylogenetically separate lineage, and Endomyxa is currently regarded as its own phylum.
The apusomonads are a group of protozoan zooflagellates that glide on surfaces, and mostly consume prokaryotes. They are of particular evolutionary interest because they appear to be the sister group to the Opisthokonts, the clade that includes both animals and fungi. Together with the Breviatea, these form the Obazoa clade.
The vampyrellids, colloquially known as vampire amoebae, are a group of free-living predatory amoebae classified as part of the lineage Endomyxa. They are distinguished from other groups of amoebae by their irregular cell shape with propensity to fuse and split like plasmodial organisms, and their life cycle with a digestive cyst stage that digests the gathered food. They appear worldwide in marine, brackish, freshwater and soil habitats. They are important predators of an enormous variety of microscopic organisms, from algae to fungi and animals. They are also known as aconchulinid amoebae.
A protist is any eukaryotic organism that is not an animal, plant, or fungus. The protists do not form a natural group, or clade, since they exclude certain eukaryotes with whom they share a common ancestor; but, like algae or invertebrates, the grouping is used for convenience. In some systems of biological classification, such as the popular five-kingdom scheme proposed by Robert Whittaker in 1969, the protists make up a kingdom called Protista, composed of "organisms which are unicellular or unicellular-colonial and which form no tissues". In the 21st century, the classification shifted toward a two-kingdom system of protists: Chromista and Protozoa.
Gromiida is an order of Endomyxa. It is the only order in the class Gromiidea.
Picophagea, also known as Synchromophyceae, is a class of photosynthetic stramenopiles. The chloroplast of the Synchromophyceae are surrounded by two membranes and arranged in a way where they share the outer pair of membranes. The entire chloroplast complex is surrounded by an additional two outer membranes.
Leucodictyids are heterotrophic amoeboid protists that comprise the order Leucodictyida in the phylum Cercozoa.
Acinetactis is a possible genus of protist first described in 1886 by A.C. Stokes, who also described the type species A. mirabilis. In 1928, Valkanov reported the discovery of a second species, A. arnaudovii. The most recently recorded specimen of A. mirabilis was reported in 1940, no Acinetactis specimens have been reported since.
Cryptofilida is an order of small heterotrophic protists in the phylum Cercozoa. They are filose amoebae that lack cilia and gliding, and are instead characterized by movement through branching or unbranched granular filopodia that are appressed to the substrate during their feeding.
Euglyphia is a group of imbricate protists in the phylum Cercozoa. They are unicellular eukaryotes characterized by a cell body covered in large imbricate scales, and an apical aperture through which they extend either filose pseudopodia or two cilia of different sizes that are not used for gliding.
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