Granofilosea

Granofilosea
Clathrulina elegans
Scientific classification
Domain:	Eukaryota
Clade:	Sar
Clade:	Rhizaria
Phylum:	Cercozoa
Subphylum:	Reticulofilosa
Class:	Granofilosea Cavalier-Smith & Bass 2009 [1]
Orders
Desmothoracida Cryptofilida Limnofilida Leucodictyida

Granofilosea is a class of cercozoan protists in the subphylum Reticulofilosa. ^[2] Out of the three groups that were traditionally considered heliozoans: the heliomonads, gymnosphaerids and desmothoracids, the latter were recently grouped into this new class.

Phylogeny and taxonomy

Phylogeny

The following cladogram shows the results of a 2011 phylogenetic analysis: ^[1]

	Monadofilosa Granofilosea novel clade Gran-5 Desmothoracida novel clade Gran-4 novel clades Gran-2 & 3 Cryptofilida novel clade Gran-1 Nanofilidae novel clade Gran-6 Mesofilidae Leucodictyida Limnofilida
	Chlorarachnea

Taxonomy

• Class Granofilosea Cavalier-Smith & Bass 2009 ^{[3] [4]}
  • Family ?Microgromiidae De Saedeleer 1934
    • Genus Apogromia de Saedeleer 1934
    • Genus Belaria de Saedeleer 1934
    • Genus Heterogromia de Saedeleer 1934
    • Genus Microgromia Hertwig & Lesser 1874
    • Genus Paralieberkuehnia de Saedeleer 1934
  • Order ?Axomonadida [Exonucleoaxoplastiales]
    • Family Tetradimorphidae Febvre-Chevalier & Febvre 1984
      • Genus Tetradimorpha Hsiung 1927
  • Order Desmothoracida Honigberg et al. 1964 [Cryptaxohelida; Desmothoraca Hertwig & Lesser 1874; Clathrulinida Starobogatov 1980]
    • Family Clathrulinidae Claus 1874
      • Genus Actinosphaeridium Zacharias 1893
      • Genus Cienkowskya Schaudinn 1896 non Regel & Rach 1859 non Solms 1867
      • Genus Clathrulina Cienkowski 1867
      • Genus Elaeorhanis Greeff 1873
      • Genus Hedriocystis Hertwig & Lesser 1874
      • Genus Penardiophrys Mikrjukov 2000
  • Order Cryptofilida Cavalier-Smith & Bass 2009
    • Family Mesofilidae Cavalier-Smith & Bass 2009
      • Genus Mesofila Cavalier-Smith & Bass 2009
    • Family Nanofilidae Cavalier-Smith & Bass 2009
      • Genus Nanofila Bass et al. 2009
  • Order Limnofilida Cavalier-Smith & Bass 2009
    • Family Limnofilidae Cavalier-Smith & Bass 2009
      • Genus Limnofila Cavalier-Smith & Bass 2009
  • Order Leucodictyida Cavalier-Smith 1993 emend. 2003
    • Family Massisteriidae Cavalier-Smith 1993
      • Genus Massisteria Larsen & Patterson 1990
      • Genus Minimassisteria Arndt & Cavalier-Smith 2011
    • Family Leucodictyidae Cavalier-Smith 1993 [Reticulamoebidae]
      • Genus Leucodictyon Grell 1991
      • Genus Reticulamoeba Grell 1994
      • Genus Thalassomyxa Grell 1985

References

1. Bass D, Chao EE, Nikolaev S, Yabuki A, Ishida K, Berney C, Pakzad U, Wylezich C, Cavalier-Smith T (2009). "Phylogeny of novel naked Filose and Reticulose Cercozoa: Granofilosea cl. n. and Proteomyxidea revised". Protist. 160 (1): 75–109. doi:10.1016/j.protis.2008.07.002. PMID   18952499.
2. Cavalier-Smith, Thomas (2017). "Kingdom Chromista and its eight phyla: a new synthesis emphasising periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences". Protoplasma. 255 (1): 297–357. doi:10.1007/s00709-017-1147-3. PMC   5756292 . PMID   28875267.
3. Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M. (2015-04-29). "A Higher Level Classification of All Living Organisms". PLOS ONE. 10 (4) e0119248. Bibcode:2015PLoSO..1019248R. doi: 10.1371/journal.pone.0119248 . ISSN   1932-6203. PMC   4418965 . PMID   25923521.
4. Adl, S.M.; Simpson, A.G.B.; Lane, C.E.; Lukeš, J.; Bass, D.; Bowser, S.S.; Brown, M.W.; Burki, F.; Dunthorn, M.; Hampl, V.; Heiss, A.; Hoppenrath, M.; Lara, E.; le Gall, L.; Lynn, D.H.; McManus, H.; Mitchell, E.A.D.; Mozley-Stanridge, S.E.; Parfrey, L.W.; Pawlowski, J.; Rueckert, S.; Shadwick, L.; Schoch, C.L.; Smirnov, A.; Spiegel, F.W. (September 2012), "The Revised Classification of Eukaryotes", Journal of Eukaryotic Microbiology, 59 (5): 429–514, doi:10.1111/j.1550-7408.2012.00644.x, PMC   3483872 , PMID   23020233

External links

• Data related to Granofilosea at Wikispecies